Allen Formation (original) (raw)

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Geological formation in Argentina

Allen Formation
Stratigraphic range: Mid Campanian-Maastrichtian~73–69 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓
Outcrop of the Allen and Anacleto Formations at Auca Mahuida
Type Geological formation
Unit of Malargüe Group
Underlies Jagüel Formation
Overlies Anacleto Formation
Thickness up to 70 m (230 ft)
Lithology
Primary Mudstone, sandstone
Other Limestone
Location
Coordinates 40°00′S 66°36′W / 40.0°S 66.6°W / -40.0; -66.6
Approximate paleocoordinates 42°48′S 52°12′W / 42.8°S 52.2°W / -42.8; -52.2
Region Neuquén, Río Negro & La Pampa Provinces
Country Argentina
Extent Neuquén Basin
Type section
Named by Uliana & Dellapé
Year defined 1981
Allen Formation is located in ArgentinaAllen FormationAllen Formation (Argentina)

The Allen Formation is a geological formation in Argentina whose strata date back to the Late Cretaceous (middle Campanian to early Maastrichtian.[1] Dinosaur remains are among the fossils that have been recovered from the formation.[2] Indeterminate chelid remains and other vertebrates have also been discovered in this formation.[3]

Fauna of Allen Formation

Uliana and Dellapé defined the formation's stratotype in 1981 in the eastern area of the Bajo de Añelo, where the relation between the base and top is clearly exposed. The deposits are mostly clastic, interbedded with banks of limestone and layers of anhydrite, which were defined as continental and shallow marine facies associated with semiarid conditions.[4]

The interpreted sedimentary paleoenvironments range from purely continental such as ephemeral lacustrine, aeolian and fluvial systems to coastal marine paleoenvironments with development of estuaries and tidal flats, followed by a lagoon sedimentary stage from marsh to sea with carbonate precipitation in an area protected from waves, ending with a retraction leading to the accumulation of evaporites.[4]

Armas and Sánchez performed a detailed facies analysis of the formation in 2015, where the authors concluded the formation represents a hybrid coastal system of tidal flats, dominated by Atlantic ingressions, with a large storm influence in some areas linked to aeolian systems.[5]

Life restoration of Dinosaurs discovered from Allen Formation

| | | | | | ------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------- | | ---------------------------------------------------------------------------------------------- | | Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon | | Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited. |

Dinosaur eggs are known from the formation.[6]

| | | | | | ------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------- | | ---------------------------------------------------------------------------------------------- | | Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon | | Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited. |

Ankylosaurs

Ankylosaurs reported from the Allen Formation
Genus Species Location Stratigraphic position Material Notes Images
Patagopelta P. cristata Salitral Moreno locality. Lower Tooth, three posterior dorsal vertebrae, caudal vertebrae, two caudal centra, right femur, partial cervical half ring, and osteoderms. A parankylosaur of uncertain affinities. [7][8]

Hadrosaurs

| | | | | | ------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------- | | ---------------------------------------------------------------------------------------------- | | Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon | | Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited. |

Hadrosaurs reported from the Allen Formation
Genus Species Location Stratigraphic position Material Notes Images
Bonapartesaurus B. rionegrensis Salitral Moreno and Islas Malvinas. Lower A partial skeleton. A hadrosaur.
Kelumapusaura K. machi Upper Partial skull, a cervical vertebrae, several sacrals, sternal rib and sternal plate, and multiple referred specimens found in a bonebed A saurolophine hadrosaur.[9]
Lapampasaurus L. cholinoi Lower Elements of the axial and appendicular skeleton of a subadult individual A hadrosaur.[10]
Willinakaqe W. salitralensis Salitral Moreno site. Lower A right premaxilla. A saurolophine hadrosaur.[11]

Sauropods

| | | | | | ------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------- | | ---------------------------------------------------------------------------------------------- | | Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon | | Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited. |

Sauropods reported from the Allen Formation
Genus Species Location Stratigraphic position Material Notes Images
Aeolosaurus Indeterminate Lower A titanosaur.[6]
Bonatitan B.reigi[12] Lower Braincases, caudal vertebrae, and several limb elements A titanosaur.[12]
Menucocelsior M. arriagadai Lower Seventeen caudal vertebrae and several appendicular bones: a right humerus, a left fibula and some metapodial. A titanosaur.
Panamericansaurus P. schroederi "Five tail vertebrata, sacral vertebrae, left humerus and rib fragments" A titanosaur.[6][14]
Pellegrinisaurus? P. powelli Lower (if it is from the formation) "Dorsal and caudal vertebrae, partial femur" A titanosaur.[6][14]
Rocasaurus R. muniozi Lower "Partial postcranial skeleton" A titanosaur.[6][15]
Theropods reported from the Allen Formation
Genus Species Location Stratigraphic position Material Notes Images
Austroraptor A. cabazai Lower A fragmentary skeleton including parts of the skull, lower jaw, a few neck and torso vertebrae, some ribs, a humerus, and assorted bones from both legs A dromaeosaurid.
Bonapartenykus B. ultimus Upper A holotype consists of a mid-dorsal vertebra, both scapulocoracoids, left tibia and femur, left pubis articulated with the pubic peduncle of the ilium, the anterior blade of the left ilium, and two partially preserved eggs. An alvarezsauroid.
Lamarqueavis L. australis Lower "Right coracoid with damaged sternal and omal extremities, and lacking acrocoracoidal process" An ornithuran.[16]
Limenavis L. patagonica Lower "Partial forelimb" An ornithuran.[6][17]
Niebla N. antiqua Upper Braincase, fragmentary jaw and teeth, relatively complete scapulocoracoid, dorsal ribs, and incomplete vertebrae. An abelisaurid[18]
Quilmesaurus Q. curriei Upper Femur and distal tibia An abelisaurid.[6][19]

Fragmentary fossils are known from the formation.[6]

Pterosaurs reported from the Allen Formation
Genus Species Location Stratigraphic position Material Notes Images
Aerotitan A. sudamericanus Bajo de Arriagada Upper Partial rostrum The first unambiguous azhdarchid from South-America[20]
Fish reported from the Allen Formation[3]
Genus Species Location Stratigraphic position Material Notes Images
Atlantoceratodus A. patagonicus 3 tooth plates A Ceratorontiform lungfish[21]
Chondrichthyes indeterminate 11 vertebral centra
Diplomystidae indeterminate 4 incomplete pectoral spines
Siluriformes indeterminate 5 incomplete pectoral spines
Lepisosteidae indeterminate 6 vertebral centra
Teleostei indeterminate 10 isolated teeth
cf. Percichthyidae indeterminate 19 fragmentary vertebrae
Squamates reported from the Allen Formation[3]
Genus Species Location Stratigraphic position Material Notes Images
Patagoniophis P. parvus incomplete trunk vertebra
Alamitophis A. argentinus incomplete trunk vertebra
Madtsoiidae indeterminate incomplete trunk vertebra
Turtles reported from the Allen Formation
Genus Species Location Stratigraphic position Material Notes Images
Yaminuechelys Y. gasperinii A chelid turtle
Rhynchocephalia reported from the Allen Formation
Genus Species Location Stratigraphic position Material Notes Images
Lamarquesaurus L. cabazai Cerro Tortuga [22]
Plesiosaurs reported from the Allen Formation
Genus Species Location Stratigraphic position Material Notes Images
Kawanectes K. lafquenianum

Unnamed frogs belonging to the family Calyptocephalellidae and Leptodactylidae, and those with no family designation were also found.[23][3]

Frogs reported from the Allen Formation
Genus Species Location Stratigraphic position Material Notes Images
Calyptocephalella C. satan calyptocephalellid frog[24]
Kuruleufenia K. xenopoides Pipid frog.[25]

The mammal fauna of the Allen Formation is known from seven teeth, which document the presence of several species.[26]

Mammals reported from the Allen Formation
Genus Species Location Material Notes Images
Mesungulatum M. lamarquensis Cerro Tortuga Two upper molars and a fragmentary lower molar A dryolestoid
Groebertherium G. stipanicici Cerro Tortuga One upper molar A dryolestoid
cf. Brandonia sp. Cerro Tortuga One lower molar A dryolestoid
Barberenia B. allenensis Cerro Tortuga One upper molariform A dryolestoid
Solanutheirum S. walshi Cerro Tortuga A right lower molar and left dentary fragment. A meridiolestid.[27]
Trapalcotherium T. matuastensis Cerro Tortuga One first lower molar A gondwanathere
Plants reported from the Allen Formation
Genus Species Location Stratigraphic position Material Notes Images
Podocarpoxylon P. mazzonii Valcheta Petrified Forest, Río Negro Province.[28] Fossil wood.[28]
  1. ^ Salgado et al., 2007
  2. ^ Weishampel, David B; et al. (2004). "Dinosaur distribution (Late Cretaceous, South America)." In: Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. Pp. 600-604. ISBN 0-520-24209-2.
  3. ^ a b c d Martinelli, A. and Forasiepi, A.M. (2004). "Late Cretaceous vertebrates from Bajo de Santa Rosa (Allen Formation), Rio Negro province, Argentina, with the description of a new sauropod dinosaur (Titanosauridae)". Revista del Museo Argentino de Ciencias Naturales 6(2): 257–305.
  4. ^ a b Armas & Sánchez, 2015, p.101
  5. ^ Armas & Sánchez, 2015, p.110
  6. ^ a b c d e f g h Weishampel et al., 2004, p.604
  7. ^ Arbour, Victoria M.; Currie, Philip J. (2016-05-03). "Systematics, phylogeny and palaeobiogeography of the ankylosaurid dinosaurs". Journal of Systematic Palaeontology. 14 (5): 385–444. Bibcode:2016JSPal..14..385A. doi:10.1080/14772019.2015.1059985. ISSN 1477-2019. S2CID 214625754.
  8. ^ Riguetti, Facundo; Pereda-Suberbiola, Xabier; Ponce, Denis; Salgado, Leonardo; Apesteguía, Sebastián; Rozadilla, Sebastián; Arbour, Victoria (2022-12-31). "A new small-bodied ankylosaurian dinosaur from the Upper Cretaceous of North Patagonia (Río Negro Province, Argentina)". Journal of Systematic Palaeontology. 20 (1): 2137441. Bibcode:2022JSPal..2037441R. doi:10.1080/14772019.2022.2137441. ISSN 1477-2019. S2CID 254212751.
  9. ^ Rozadilla, S.; Brissón-Egli, F.; Agnolín, F.L.; Aranciaga-Rolando, A.M.; Novas, F.E. (2022). "A new hadrosaurid (Dinosauria: Ornithischia) from the Late Cretaceous of northern Patagonia and the radiation of South American hadrosaurids". Journal of Systematic Palaeontology. 19 (17): 1207–1235. doi:10.1080/14772019.2021.2020917. S2CID 247122005.
  10. ^ Coria et al., 2012
  11. ^ Juárez Valieri et al., 2010
  12. ^ a b "63.10 Provincia de Río Negro, Argentina; 4. Río Colorado Formation," in Weishampel, et al. (2004). Page 604.
  13. ^ a b "Table 13.1," in Weishampel, et al., 2004, p.269
  14. ^ "Table 13.1," in Weishampel, et al., 2004, p.270
  15. ^ Agnolin, 2010
  16. ^ "Table 11.1," in Weishampel, et al., 2004, p.215
  17. ^ Aranciaga Rolando, Mauro; Cerroni, Mauricio A.; Garcia Marsà, Jordi A.; Agnolín, Federico l.; Motta, Matías J.; Rozadilla, Sebastián; Brisson Eglí, Federico; Novas, Fernando E. (2020-10-14). "A new medium-sized abelisaurid (Theropoda, Dinosauria) from the late cretaceous (Maastrichtian) Allen Formation of Northern Patagonia, Argentina". Journal of South American Earth Sciences. 105: 102915. doi:10.1016/j.jsames.2020.102915. hdl:11336/150468. ISSN 0895-9811. S2CID 225123133.
  18. ^ "Table 4.1," in Weishampel, et al., 2004, p.77
  19. ^ Novas et al., 2012
  20. ^ Federico Agnolin (2010). "A new species of the genus Atlantoceratodus (Dipnoiformes: Ceratodontoidei) from the Uppermost Cretaceous of Patagonia and a brief overview of fossil dipnoans from the Cretaceous and Paleogene of South America". Brazilian Geographical Journal: Geosciences and Humanities Research Medium. 1 (2): 162–210.
  21. ^ Apesteguía & Rougier, 2007, p.3
  22. ^ Suazo Lara, F.; Gómez, R. O. (2022-03-01). "In the shadow of dinosaurs: Late Cretaceous frogs are distinct components of a widespread tetrapod assemblage across Argentinean and Chilean Patagonia". Cretaceous Research. 131: 105085. Bibcode:2022CrRes.13105085S. doi:10.1016/j.cretres.2021.105085. ISSN 0195-6671. S2CID 243482426.
  23. ^ Federico Agnolin (2012). "A new Calyptocephalellidae (Anura, Neobatrachia) from the Upper Cretaceous of Patagonia, Argentina, with comments on its systematic position". Studia Geologica Salmanticensia. 48 (2): 129–178.
  24. ^ Gómez, Raúl O. (2016-07-01). "A new pipid frog from the Upper Cretaceous of Patagonia and early evolution of crown-group Pipidae". Cretaceous Research. 62: 52–64. Bibcode:2016CrRes..62...52G. doi:10.1016/j.cretres.2016.02.006. hdl:11336/59544. ISSN 0195-6671.
  25. ^ Rougier et al., 2009
  26. ^ Connelly, B. E.; Cardozo, M. S.; Montgomery, J. D.; Rougier, G. W. (2024). "New mammals from the Upper Cretaceous Allen Formation (Patagonia, Argentina) and reassessment of meridiolestidan diversity". Cretaceous Research. 162. 105935. doi:10.1016/j.cretres.2024.105935.
  27. ^ a b Passalia, M. G.; Garrido, A.; Iglesias, A.; Vera, E. I. (2022-10-14). "The Valcheta Petrified Forest (Upper Cretaceous), Northern Patagonia, Argentina: A Geological And Paleobotanical Survey". Cretaceous Research. 142: 105395. doi:10.1016/j.cretres.2022.105395. ISSN 0195-6671. S2CID 252964524.