Neuroptera (original) (raw)

After Palker, 1982, Grimaldi & Engel, 2005, and Engel & Grimaldi, 2007

<==o Neuroptera Linnaeus sensu stricto? [Planipennia] (verkkosiipiset (harsosiipiset); lacewings, antlions, owflies, etc.) |?- †Mesohemerobius jeholensis Ping, 1928; L.Cret. EAs. [Neuroptera incertae sedis (Makarkin et al., 2003)] |-- †“Permithonidae” --+--o Nevrorthidae Nakahara [Neurorthidae (_lapsus calami_)] [Nevorthiformia] | |-- †_Rophalis relicta_ (Hagen); M.Eoc. NEu. | |-- _Austroneurorthus_ | |--o _Nevorthus_ | | -- N. fallax | -- _Nipponeurorthus_ --+?- †Grammosmylidae |--o Hemerobiiformia (harsokorentomaiset) | |?- †“Osmylitidae” | --+?-o †_Prohemerobiidae_ | | -- †Prohemerobius Handlirsch, 1906 | |?- †“Archaeosmylus_” complesus Whalley, 1988 | |?- †“_Archaeosmylus_” alysius Whalley, 1988 | |--+--o Ithonoidea | | | |-- Polystoechotidae Handlirch 1906 (giant lacewings) | | | -- [Ithonidae](ithonidae.html) Newman, 1853 [incl. Rapismatidae Krüger, 1922] (moth lacewings) | | --+-- Chrysopidae Schneider (green lacewings; harsokorennot) | | --o Osmyloidea | | |**==**o †Archeosmylidae Riek, 1953 [probably paraphyletic basal Osmyloids] | | | |--o †_Archaeosmylus_ Riek, 1953; L.Perm.-E.Jur. Aust. | | | | |-- †_A. pectinatus_ Reik, 1953 | | | | |?- †_A. stigmatus_ Reik, 1955 | | | | |?- †_A. costalis_ Reik, 1955 | | | | ?- †_A. sp. (L.Trias. Aust) | | | |?- †Archeosmylidae gen. et sp. indet. (L.Perm. South Africa) | | | |-- †Babykamenia eskovi Ponomarenko & Shcherbakov, 2004 | | | ?-o †_Lithosmylidia_ Reik, 1955; M.-L.Trias. Aust. | | | |-- †_L. lineata_ Reik, 1955 | | | |?- †_L. barrone_ Lambkin, 1988 | | | ?- †L. parvula Lambkin, 1988 | | --o Osmylidae Leach, 1815 (osmylids) | | |-- †_Archaeosmylidia fusca_ Makarkin, Yang & Ren, 2012; M.Jur. EAs. | | --+?- †Tetanoptilon Bode, 1953 | | |-- †Osmalochrysa Jepson, Makarkin & Coram, 2012 | | |?-o †Mesosmylininae | | | |-- †Mesosmylina Bode, 1953 | | | |-- †Sogjuta Martynova, 1958 | | | -- †_Petrushevskia_ Martynova, 1958 | | |?-o †Saucrosmylinae Ren & Yin, 2003 | | | |-- †_Saucrosmylus sambneurus_ Ren & Yin, 2003; M.Jur. EAs. | | | |-- †_Rudiosmylus ningchengensis_ Ren & Yin, 2003; M.Jur. EAs. | | | |-- †_Laccosmylus calophlebius_ Ren & Yin, 2003; M.Jur. EAs. | | | --o †Bellinympha Y. Wang, Ren, Liu, Shih & Engel, 2010 in Y. Wang, Liu, X. Wang, Shih, Zhao, Engel & Ren, 2010; M.Jur. EAs. | | | |-- †B. filicifolia Y. Wang, Ren, Liu, Shih & Engel, 2010 in Y. Wang, Liu, X. Wang, Shih, Zhao, Engel & Ren, 2010 | | | -- †_B. dancei_ Y. Wang, Ren, Liu, Shih & Engel, 2010 _in_ Y. Wang, Liu, X. Wang, Shih, Zhao, Engel & Ren, 2010 | | |--o Protosmylinae | | | |-- _Gryposmylus_ | | | |-- _Heterosmylus_ | | | |-- _Paryphosmylus_ | | | |-- †_Juraheterosmylus_ | | | |-- †_Mesosmylus_ | | | |-- †_Osmylidia_ | | | |-- †_Protosmylina_ | | | --o †Protosmylus | | | -- †_P. pictus_ (Hagen); M.Eoc. NEu. | | |--o Gumillinae | | |--o Spilosmylinae | | | -- Thyridosmylus | | |-- Kempyninae | | |-- Porismus strigatus (Burmeister, 1839) [Porisminae] | | |-- Stenosmylinae | | |-- Eidoporisminae | | --o Osmylinae | | --o Osmylus | | -- _O. fulvicephalus_ | --+-- Hemerobiidae Latreille [Hemerobioidea] (brown lacewings; kirvakorennot) | --+--+--o Sisyridae (spongillaflies; rantakorennot) | | | |?- †_Cratosisyrops_ L.Cret. ESA. (_incertae sedis_) | | | |?- †_Sisyra_? _amissa_ Hagen; M.Eoc. NEu. | | | |**==** numerous **Eocene** fossil genera | | | |?- _Climacia_ | | | |?- _Sisyra_ | | | == numerous Recent genera | | -- [Coniopterygidae](coniopterygidae.html) Burmeister [Coniopterygoidea] (dustywings; vahakorennot) | --o Mantispoidea | |--o Dilaridae Newman (pleasing lacewings) | | |--o Nallachiinae | | | --o _Nallachius_ | | | -- N. americanus | | --o Dilarinae | | |-- †_Cascadilar eocenicus_ Engel, 1999; M.Eoc. NEu. | | --o Dilar | | -- _D. septentrionalis_ | --+-- Mantispidae Leach (mantispids; rukoilijakorennot) | --+-- †“Mesithonidae” | --+--o Rhachiberothidae Tjeder (thorny lacewings) | | |-- †Paraberotha acra Whalley; L.Cret. SWAs. | | |-- †Retinoberotha stuermeri Schlüter; U.Cret. WEu. | | |-- †Eorhachiberotha burmitica Engel, 2004; U.Cret. SEAs. | | |-- †Rhachibermissa splendida Grimaldi; U.Cret. ENAm. | | |-- †Whalfera venatrix (Whalley; 1983) M.Eoc. NEu. | | |-- †Alloberotha petrulevicii Nel et al., 2005; U.Cret. WEu. | | |-- †Chimerhachiberotha acrasarii Nel et al., 2005; L.Cret. SWAs. | | |-- †Spinoberotha mickaelacrai Nel et al., 2005; L.Cret. SWAs. | | |-- †Oisea celinea (Nel et al., 2005) Eoc. WEu. | | **==** **Recent genera** | --o Berothidae Handlirsch (beaded lacewings) | |?- †Banoberotha enigmatica Whalley; U.Cret. SWAs.. | |?-o †Jersiberotha Grimaldi; U.Cret. NAm. | | |-- †J. luzzii Grimaldi | | -- †_J. similis_ Grimaldi | |?- †_Nascimberotha picta_ Grimaldi; U.Cret. NAm. | |?- †_Microberotha macculloughi_ Archibald & Makarkin; Eoc. | |?- †_Plesiorobius_? _anadensis_ Klimaszewski & Kevan; U.Cret. NAm. | |?- †_Proberotha prisca_ Krüger, 1923; M.Eoc. NEu. | |?- _Lomamyia_ | |-- Cyrenoberothinae | |-- Berothinae | |-- Nosybinae | |-- Nyrminae | |-- Protobiellinae | -- Trichomatinae --o Myrmeleontiformia (muurahaiskorentomaiset) |?- †_Aetheogramma speciosa_ Ren & Engel, 2008 [Aetheogrammatidae Ren & Engel, 2008] --o Myrmeleontoidea |--o Nemopteroidea | |--o †Kalligrammatidae Handlirsch, 1906 | | |--o †Kalligrammatinae Handlirsch, 1906 | | | |-- †Huiyingogramma formosum Liu et al., 2014 | | | |--o †Kalligramma Walther, 1904 | | | | -- †_K. paradoxum_ Liu et al., 2014 | | | |--o †_Limnogramma_ Ren, 2003 | | | | |-- †_L. hani_ | | | | |-� †_L. mira_ | | | | -- &38224;L. mongolicum | | | |--o †Meioneurites Handlirsch, 1906; L.Jur. CAs. | | | | |-- †M. (Parameioneurites) spectabilis Engel, 2005 | | | | |-- †M. schlosseri Handlirsch, 1906 | | | | -- †_M. villosus_ Panfilov, 1980 | | | |-- †_Protokalligramma bifasciatum_ Yang, Makarkin & Ren, 2011 | | | --o †Sophogramma Ren & Guo, 1996 | | | |-- †S. papilionaceum Ren & Guo, 1996 | | | -- †_S. wimbledoni_ Jepson, Makarkin & Coram, 2012 | | --o †Kallihemerobiinae Ren & Engel, 2008 | | |-- †Apochrysogramma rotundum Yang, Makarkin & Ren, 2011 | | -� †_Kallihemerobius pleioneurus_ Ren & Oswald, 2002 | --+-- Psychopsidae Handlirsch (silky lacewings) | -- [Nemopteridae](nemopteridae.html) (spoon-, thread-winged lacewings; lippukorennot) --+--o †Osmylopsychopidae | -- †_Petropsychops superba_ Trias. Aust. |-- †Nymphitidae |-- †“Solenoptilidae” |-- †Brogniartiellidae --+--o Nymphidae Rambur [incl. Myiodactylidae] (split-footed lacewings) | |-- †Pronymphes mengeanus (Hagen); M.Eoc. NEu. | |-- Nymphes | |-- Myiodactylus | |-- Osmylops | **==** numerous other Recent genera --+-- †Babinskaiidae |--o Myrmeleontidae Latreille [incl. †Palaeoleontidae] (antlions; muurahaiskorennot, -leijonat) | |?- †Samsonileon fragmentus Ponomarenko; L.Cret. SWAs. [†Palaeoleontidae] | |?- †Palaeoleon ferrogeneticus Rice; U.Cret. NENAm. [†Palaeoleontidae] | |?- †Dendroleon septemmontanus Statz, 1936 | |?-o Porrelus | | |-- †P. dominicanus Poinar & Stange, 1996; Mioc. CAm. | | **==** **recent _Porrelus_ -species** | |?- Dimarinae | |?- Stilbopteryginae | |?- Pseudimarinae | |?- Echthromyrmicinae | |?- Palparinae [incl. †Araripeneurinae] | |?- Myrmecaelurinae | |?- Acanthaclisinae | |?- Myrmeleontinae | |?- Brachynemurinae | |?- Dendroleontinae | |?- Glenurinae | ?- Nemoleontinae --o Ascalaphidae Rambur (owlflies; pöllökorennot) |?- †_Ascaloptynx oligocenicus_ Nel, 1990; Olig. WEu. |?- †_Borgia proavus_ (Hagen, 1858) Navás, 1913; Olig. CEu. |?- †_Mesascalaphus yangi_ Ren et al., 1995; U.Jur. EAs. |?- †_Neadelphus protae_ MacLeod, 1970; M.Eoc. NEu. [larva] |?- †_Prosuhpalacsa biamoensis_ Makarkin, 1998; ?U.Olig.-?L.Mioc. EAs. |?- †_Ricartus edwardsi_ (Oustalet, 1870) Navás, 1913; Olig. WEu. |?- _Cordulecerus_ |?- Albardiinae |--o Ascalaphinae | |--o _Ululodes_ | | -- †U. paleonesia Engel & Grimaldi, 2007; Mioc. CAm. | **==** **Other Ascalaphines** --o Haplogleniinae |--o Amoea | -- †_A. electrodominicana_ Engel & Grimaldi, 2007; Mioc. CAm. == Other Haplogleniines

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