Rhynchosauria (original) (raw)

After Langer & Schultz, 2000, Hone & Benton, 2008, Langer, Montefeltro, Hone, Whatley & Schultz, 2010, and Ezcurra, Montefeltro & Butler, 2016

**<==**o †Rhynchosauria Osborn, 1903 (rhynchosaurs; rynkosaurit) |?- †Noteosuchus colletti (Watson, 1912) Broom, 1925 [Eosuchus colletti Watson, 1912] [preoccupied by Eosuchus lerichei_ Dollo, 1907, a gavialoid crocodylian] |?- †_Colobops noviportensis Pritchard, Gauthier, Hanson, Bever & Bhullar, 2018 |-- †Mesosuchus browni Watson, 1912; E.Trias. SAf. [“Mesosuchinae”] --+-- †_Howesia browni_ Broom, 1905; E.Trias. SAf. --+-- †Eohyosaurus wolvaardti Butler, Ezcurra, Montefeltro, Samathi & Sobral, 2015; M.Trias. SAf. --o †Rhynchosauridae _sensu_ Butler et al., 2015 [incl. Hyperdapedontidae Lydekker, 1885] |?- †_Eifelosaurus triadicus_ Jaekel, 1904; E.Trias. Eu. |?- †_Mesodapedon kuttyi_ Chatterjee, 1980; L.Trias. SAs. |--o †Rhynchosaurinae | -- †Rhynchosaurus articeps Owen, 1841; M.-L.Trias. --o †Hyperdapedontidae (Lydekker, 1885) Langer, Montefeltro, Hone, Whatley & Schultz, 2010 [Hyperdapedontinae Chatterjee, 1969] --+-- †Langeronyx brodiei (Benton, 1990) Ezcurra, Montefeltro & Butler, 2016 [Rhynchosaurus brodiei Benton, 1990] --+--o †Stenaulorhynchinae _sensu_ Ezcurra, Montefeltro & Butler, 2016 | |-- †_Stenaulorhynchys stockleyi_ (Haughton, 1932) Huene, 1938 [_Hyperodepedon stockleyi_; _Scaphonyx stockleyi_; _Supradapedon stockleyi_] M.Trias. EAf. | -- “Mariante rhynchosaur” --+-- †_Bentonyx sidensis_ Langer, Montefeltro, Hone, Whatley & Schultz, 2010 --+-- †Fodonyx spenceri (Benton, 1990) Hone & Benton, 2008 [Rhynchosaurus spenceri Benton, 1990] --+-- †_Ammorhynchus navajoi_ Nesbitt & Whatley, 2004; M.Trias. SWNAm. --o †Hyperdapedontidae (Lydekker, 1885) Langer, Montefeltro, Hone, Whatley & Schultz, 2010 |-- †Isalorhynchus genovefae Buffetaut, 1983 [=? Hyperodepedon Huxley, 1859 (Langer et al., 2000] --o †Hyperdapedontinae (Lydekker, 1885) Chatterjee, 1969 _sensu_ Langer & Schultz, 2000 |-- †_Teyumbaita sulcognathus_ (Azevedo & Schultz, 1987) Montefeltro, Langer & Schultz, 2010 [“Scaphonyx” _sulcognathus_ Azevedo & Schultz, 1987] |-- †_Otischalia elderae_ Hunt & Lucas, 1991 --o †Hyperodepedon Huxley, 1859 [Stenometopon Boulenger, 1903; Scaphonyx Woodward, 1907; Macrocephalosaurus Tupi-Caldas, 1933; Paradapedon Huene, 1938; Cephalonia Huene, 1942; Supradapedon Chatterjee, 1980] L.Trias. Eu. Ind. SAs. SAm. |?- †H. australis (Huene, 1926) [Scaphonyx australis Huene, 1926] (nomen dubium?) |?- †H. lotziana (Huene, 1942) [Cephalonia lotziana Huene, 1942] (nomen dubium?) |-- †H. huenei Langer & Schultz, 2000 [Scaphonyx fischeri Woodward, 1907 in partim] L.Trias. SAm. --+-- †_H. mariensis_ (Tupi-Caldas, 1933) L.Trias. SAm. |--+-- †_H. huxleyi_ Lydekker, 1881 [_Paradapedon huxleyi_ (Lydekker, 1881) Huene, 1938], L.Trias. Ind. | -- †H. gordoni Huxley, 1859, L.Trias. WEu. --+?- †_H. fischeri_ (Woodward, 1907) _sensu_ Langer & Schultz, 2000 [_Scaphonyx fischeri_ Woodward, 1907; _Cephalonia_; _Cephalastron_; _Cephalastronius_; _Scaphonychimus_] M. Trias. SA. [_nomen dubium_?] -- †H. sanjuanensis (Sill, 1970) sensu Langer & Schultz, 2000 [Scaphonyx sanjuanensis Sill, 1970]

| After Ezcurra, Montefeltro & Butler, 2016 **<==**o †Rhynchosauria |-- †Noteosuchus colletti | -- †Mesosuchus browni | -- †Howesia browni | -- †Eohyosaurus wolvaardti `--o †Rhynchosauridae | -- †Rhynchosaurus articeps_ `--+--o †Stenaulorhynchinae | | -- “Mariante rhynchosaur” | `-- †Stenaulorhynchys stockleyi `--+-- †_Langeronyx brodiei `--+-- †Bentonyx sidensis_ `--+-- †_Fodonyx spenceri `--o †Hyperdapedontinae | -- †Isalorhynchus genovefae_ `--+-- †_Teyumbaita sulcognathus `-- †Hyperodepedon_ Reference(s): Ezcurra, M. D., Montefeltro, F. & Butler, R. J., 2016: The early evolution of rhynchosaurs.–Frontiers in Ecology and Evolution: Vol. 3, #142 [**doi: 10.3389/fevo.2015.00142**] | After Langer, Montefeltro, Hone, Whatley & Schultz, 2010 **<==**o †Rhynchosauria |-- †Rhynchosaurus articeps `--o †Hyperdapedontidae | --+-- “Mariante rhynchosaur” | `-- †Stenaulorhynchys stockleyi | -- †Bentonyx sidensis `--+-- †_Fodonyx spenceri `--o †Hyperdapedontinae | -- †Isalorhynchus genovefae_ `--+-- †_Teyumbaita sulcognathus `-- †Hyperodepedon Reference(s): Langer, M. C., Montefeltro, F. C., Hone, D. E., Whatley, R. & Schultz, C. L., 2010: On Fodonyx spenceri and a New Rhynchosaur from the Middle Triassic of Devon.–Journal of Vertebrate Paleontology: Vol. 30, #6, pp. 1884-1888 [**doi: 10.1080/02724634.2010.521901**] | | ------------------------------------------------------------------------------------------------- | ----------------------- | -------------------- | ------------------------------------------------------ | --------------------------------------------------------------- | | ------------------------- | -------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------- | -------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------- | ---------------------------------------------------------------------------------------------------------------------------------------------- | ---------------------------- | ---------------------------------- | ------------------------------------------------------------------------------------------- | ------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------- |

Reference(s):

• Butler, R. J., Ezcurra, M. D., Montefeltro, F. C., Samathi, A. & Sobral, G., 2015: A new species of basal rhynchosaur (Diapsida: Archosauromorpha) from the early Middle Triassic of South Africa, and the early evolution of Rhynchosauria.
  –Zoological Journal of the Linnean Society: Vol. 174, #3, pp. 571-588 [**doi: 10.1111/zoj.12246**]
• Carroll, R. L., 1988: Vertebrate paleontology and evolution.
  –W. H. Freeman and company, New York, 1988, 698.
• Carroll, R. L., 1988: Appendix. 594-648.
  in Carroll, R. L. 1988: Vertebrate paleontology and evolution.
  –W. H. Freeman and company, New York, 1988, 698
• Ezcurra, M. D., Montefeltro, F. & Butler, R. J., 2016: The early evolution of rhynchosaurs.
  –Frontiers in Ecology and Evolution: Vol. 3, #142 [**doi: 10.3389/fevo.2015.00142**]
• Ezcurra, M. D., Trotteyn, M. J., Fiorelli, L. E., Baczko, M. B., Taborda, J. R. A., Iberlucea, M. & Desojo, J. B., 2013: The oldest rhynchosaur from Argentina: a Middle Triassic rhynchosaurid from the Cha�ares Formation (Ischigualasto�Villa Uni�n Basin, La Rioja Province).
  –Pal�ontologische Zeitschrift: in press pp. 1-8 [**doi: 10.1007/s12542-013-0203-3**]
• Hone, D. W. E. & Benton, M. J., 2008: A new genus of rhynchosaur from the Middle Triassic of south-west England.
  –Palaeontology: Vol. 51, #1, pp. 95-115 [**doi: 10.1111/j.1475-4983.2007.00739.x**]
• Langer, M. C., Montefeltro, F. C., Hone, D. E., Whatley, R. & Schultz, C. L., 2010: On Fodonyx spenceri and a New Rhynchosaur from the Middle Triassic of Devon.
  –Journal of Vertebrate Paleontology: Vol. 30, #6, pp. 1884-1888 [**doi: 10.1080/02724634.2010.521901**]
• Langer, M. C. & Schultz, C. L., 2000: A new species of the Late Triassic rhynchosaur Hyperodapedon from the Santa maria Formation of South Brazil.
  –Paleontology: Vol. 43, #4, pp. 633-652
• Montefeltro, F. C., Langer, M. C. & Schultz, C. L., 2010: Cranial anatomy of a new genus of hyperodapedontine rhynchosaur (Diapsida, Archosauromorpha) from the Upper Triassic of southern Brazil.
  –Earth and Environmental Science Transactions of the Royal Society of Edinburgh: Vol. 101, #01, pp. 27-52 [**doi: doi:10.1017/S1755691010009060**]
• Nesbitt, S. J. & Whatley, R. L., 2004: The first discovery of a rhynchosaur from the upper Moenkopi Formation (Middle Triassic) of northern Arizona.
  –PaleoBios: Vol. 24, #3, pp. 1-10
• Pritchard, A. C., Gauthier, J. A., Hanson, M., Bever, G. S. & Bhullar, B.-A. S., 2018: A tiny Triassic saurian from Connecticut and the early evolution of the diapsid feeding apparatus.
  –Nature Communications: Vol. 9, #1, Art.nr. 1213, pp. (1-10) [**doi: 10.1038/s41467-018-03508-1**]