Anguidae: Glyptosaurinae (original) (raw)

After Sullivan, 1986, Zonneveld, Gunnell & Bartels, 2000, Augé, 2003, Sullivan & Augé, 2006, and Augé & Sullivan 2006

<==o †Glyptosaurinae Marsh, 1872 sensu Sullivan, 1986 |?- †Paraxestops stehlini Hoffstetter, 1962; U.Eoc. Eu. |--o †Odaxosaurus [†Pancelosaurus Meszoely 1970] [‘Odaxosaurini’ Sullivan 1979] | |-- †O. jepseni Meszoley et al., 1978 | |-- †O. niger Gilmore, 1928 | |-- †O. piger Gilmore, 1928 [Odaxosaurus obliquus Gilmore 1928; Pancelosaurus piger; Peltosaurus piger] | -- †_O. priscus_ Gao & Fox, 1996 --o †‘Melanosaurini’ Sullivan 1979 [prob. paraphyletic (Augé & Sullivan, 2006)] |-- †Proxestops silberlingii (Gilmore, 1938) Gauthier, 1982 [P. jepseni Gauthier, 1982] --+?- †_Xestops vagans_ (Marsh, 1873) Cope, 1873 _sensu_ Keller et al., 1991 [_Oreosaurus_] Eoc. NA. |?-o †_Peltosaurus_ Cope, 1872 | |?- †_P. minimus_ | |-- †_P. floridanus_ Vanzolini, 1952 | |-- †_P. granulosus_ Cope, 1873 [_Peltosaurus abbotti_ Gilmore, 1928] | -- †P. macrodon Brattstrom, 1955 |--+-- †Arpadosaurus gazinorum Meszoely, 1970; Eoc. CEu. | |-- †Melanosaurus maximus Gilmore, 1928; Eoc. NAm. | |-- †Paraplacosauriops quercyi (Filhol, 1882) Augé & Sullivan, 2006 [†Plestiodon quercyi Filhol, 1882; Diploglossus cadurcensis De Stefano, 1903; Placosaurus leenhardti Leenhardtm 1926; Placosaurus quercyi Hoffstetter, 1944; Placosaurus quercyi (Filhol, 1882) Sullivan & Augé, 2006] | --o †_Placosauriops_ Kuhn, 1940; Eoc. Eu. [**[taxonomic history](history%5FPlacosauriops.html)**] | |-- †_P. weigelti_ (Kuhn, 1940) [**[taxonomic history](history%5FPlacosauriops.html#P%5Fweigelti)**] | ?- ?†Plecosauriops indet. (Augé, 2003) --o †Glyptosaurini Sullivan, 1979 |?- †_Proglyptosaurus huerfanensis_ [_Eoglyptosaurus donohoei_ Sullivan, 1979 (_in partim_)] [**AMNH 7431**] |--+--o †_Glyptosaurus_ Marsh, 1871 [**[taxonomic history](history%5FGlyptosaurus.html)**] | | |-- †_G. anceps_ Marsh, 1871 | | -- †G. sylvestris Marsh, 1871 [**taxonomic history] | ?-o †_Paraglyptosaurus_ Sullivan, 1979 [=? _Glyptosaurus_ (Sullivan, 1986)] | |?- †_P. yatkolai_ Sullivan, 1979 | |-- †_P princeps_ (Marsh, 1872) [= _Glyptosaurus sylvestris_ Marsh, 1871 (_sensu_ Sullivan, 1986)] | -- †P. hillsi Gilmore, 1928 --+-- †_Helodermoides tuberculatus_ Douglass, 1903 --o †Placosaurus Gervais, 1848-52 [**taxonomic history] |?- †P. cayluxi (nomen dubium) |?- †P. galliae (nomen dubium) |?- †P. waltheri (nomen dubium) |?- †P. margariticeps (Gervais, 1876) (nomen dubium, type lost) [taxonomic history] |?- †P.? europaeus (Fihol, 1876) [taxonomic history] |-- †P. rugosus Gervais, 1848-52 |-- †P. mongoliensis (Sullivan, 1979) [taxonomic history] `-- †P. estesi Sullivan & Augé, 2006

| After Conrad & Norell, 2008 (fig. 10b) **<==**o †Glyptosaurinae |-- †Odaxosaurus `--+--+-- †Xestops_ | `-- †_Proxestops `--+--+-- †Paraplacosauriops_ | `-- †_Peltosaurus `--+--o †Melanosaurini | | -- †Arpadosaurus_ | `-- †_Melanosaurus `--o †Glyptosaurini | -- †Placosaurus rugosus_ | -- †Glyptosaurus `--+-- †_Proglyptosaurus huerfanensis `--+-- †Paraglyptosaurus_ `--+-- †_Helodermoides `--+-- †Placosaurus mongoliensis_ `-- †_Placosaurus estesi Reference(s): Conrad, J. L. & Norell, M. A., 2008: The braincases of two glyptosaurines (anguidae, Squamata) and anguid phylogeny.–American Museum Novitates: #3613, pp. 1-24 [**doi: 10.1206/586.1**] | After Sullivan, 1986 **<==**o †Glyptosaurinae |-- †Odaxosaurus `--+-- **Glyptosaurinae** sp. et gen. indet. `--+-- †Proxestops `--+?- †“Xestops_” | ?- †Peltosaurus | --+-- †Arpadosaurus | `-- †_Melanosaurus `--+--+-- †Helodermoides_ | `-- †_Placosaurus `--+-- †Eoglyptosaurus_ `--+-- †_Glyptosaurus `-- †Paraglyptosaurus Reference(s): Sullivan, R. M., 1986: The skull of Glyptosaurus sylvestris Marsh, 1871 (Lacertilia: Anguidae).–Journal of Vertebrate Paleontology: Vol. 6, #1, pp. 28-37 | | --------------------------------------------------------------------------------------------------------------- | -------------------------------------------------------- | ---------------------------------------------------- | | ------------------ | ----------------------------------------------------- | ------------------------- | ----------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------- | ------------------------------------------------------------------------------------------------------------------------------------------------------------------------------- | ----------------- | --------------------- | ------------------------------------------------------------ | ------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------ |

Reference(s):

• Augé, M., 2003: La faune de Lacertilia (Reptilia, Squamata) de l'Éocène inférieur de Prémontré (Bassin de Paris, France)
  [The lizard fauna (Reptilia, Squamata) from the lower Eocene locality of Prémontré (Paris Basin, France)].
  –Geodiversitas: Vol. 25, #3, pp. 539-574
• Augé, M. & Sullivan, R. M., 2006: A new genus, Paraplacosauriops (Squamata, Anguidae, Glyptosaurinae), from the Eocene of France.
  –Journal of Vertebrate Paleontology: Vol. 26, #1, pp. 133-137
• Carroll, R. L., 1988: Appendix. 594-648.
  in Carroll, R. L. 1988: Vertebrate paleontology and evolution.
  –W. H. Freeman and company, New York, 1988, 698
• Conrad, J. L. & Norell, M. A., 2008: The braincases of two glyptosaurines (anguidae, Squamata) and anguid phylogeny.
  –American Museum Novitates: #3613, pp. 1-24 [**doi: 10.1206/586.1**]
• Sullivan, R. M., 1979: Revision of the Paleogene genus Glyptosaurus (Reptilia, Anguidae).
  –Bulletin of the American Museum of Natural History: #163, pp. 1-72
• Sullivan, R. M., 1986: The skull of Glyptosaurus sylvestris Marsh, 1871 (Lacertilia: Anguidae).
  –Journal of Vertebrate Paleontology: Vol. 6, #1, pp. 28-37
• Sullivan, R. M., 1989: Proglyptosaurus huerfanensis, new genus, new species: Glyptosaurine lizard (Squamata. Anguidae) from the Early Eocene of Colorado.
  –American Museum Novitates: #2949, pp. 1-8
• Sullivan, R. M. & Augé, M., 2006: Redescription of the holotype of Placosaurus rugosus Gervais 1848-1852 (Squamata, Anguidae, Glyptosaurinae) from the Eocene of France and a revision of the genus.
  –Journal of Vertebrate Paleontology: Vol. 26, #1, pp. 127-132
• Zonneveld, J.-P., Gunnell, G. F. & Bartels, W. S., 2000: Early Eocene fossil vertebrates from the southwestern Green River basin, Lincoln and Uinta Counties, Wyoming.
  –Journal of Vertebrate Paleontology: Vol. 20, #2, pp. 369-386