Scincomorpha (original) (raw)

After Rieppel, 1994, Evans & Barbadillo, 1997, Evans & Chure, 1998, Gao & Norell, 2000, and Alifanov, 2000

Alternative phylogeny of Scincomorpha after Kearney, 2003

<==o Scincomorpha Camp, 1923 (skink-like lizards; skinkkimäiset liskot) |?- †Scincomorpha genera et species indet. [Ceder Mountain formation, Utah, USA.] [Nydam, 2002] |?- †Scincomorpha genera et species indet. [Ukhaa Tolgod, Djadokhta formation, Mongolia] [Gao & Norell, 2000] |?- †Calanguban alamoi Sim�es, Caldwell & Kellner, 2014; E.Cret. SAm. |?- †Bothriagenys mysterion Nydam, 2002 |?- †Globaura venusta Borsuk-Bialynicka, 1988 |-- †Tepexisaurus tepexii Reynoso & Callison, 2000 --+--+?- †_Changetisaurus estesi_ Federov & Nessov, 1992 [_Chengetisaurus_ lapsus calami] [Dorsetisauridae: Changetisaurinae Federov & Nessov, 1992] | --o †Paramacellodidae Estes, 1983 [a paraphyletic form-family?] | |?- †Atokasaurus metarsiodon Nydam & Cifelli, 2002b, L. Cret. NA. [cf. “Paramacellodidae”] | |?- †Pseudosaurillus becklesi Hoffstetter, 1967 | |?-o †Saurillus Owen, 1855 | | |-- †S. henkeli Hoffstetter, 1967 | | |-- †S. obtusus Owen, 1855 | | -- †_S. robustidens_ Hoffstetter, 1967 | |?- †_Saurillodon proraformis_ Estes, 1983 | |--o †_Paramacellodus_ Hoffstetter, 1967; U. Jur. L. Cret. NA. | | |?- cf. _P. sp._ [Ceder Mountain formation, Utah, USA.] [Nydam, 2002] | | |?- cf. †_P. keebleri_ Nydam & Cifelli, 2002b, L. Cret. NA. | | |-- †_P. oweni_ Hoffstetter, 1967 U. Jur. NA. | | -- †P. keebleri Nydam & Cifelli, 2002b | |--o †Becklesius Estes, 1983 | | |-- †B. hoffstetteri (Seiffert, 1973) [Macellodus brodei Hoffsteetr, 1967 (non Macellodus brodei Owen, 1854); Becklesisaurus hoffstetteri Seiffert, 1973] | | |-- †B. cataphractus Richter, 1994 | | |-- †B. sp. [Richter, 1994] | | -- †_B. nopcsai_ Folie & Codrea, 2005 | |-- †_Sharovisaurus karatauensis_ Hecht & Hecht, 1984; U. Jur. CAs. | -- †Mimobecklesiosaurus gansuensis Li, 1985; U. Jur. EAs. |--o Scincoidea Oppel, 1811 (skink-related lizards; skinkinsukuiset liskot) | |?- †Dimekodontosaurus madseni Nydam, 2002 | |?- †Hymenosaurus clarki Gao & Norell, 2000 | |?- †Scincoideus haininensis Folie, Sig� & Smith, 2005, Pal. Eu. | |?- †Parmeosaurus scutatus Gao & Norell, 2000 | |?-o †Slavoiidae Alifanov, 1993 | | |-- †Eoxanta lacertifrons Borsuk-Bialynicka, 1988 | | -- †_Slavoia darevskii_ Sulimski, 1984 | |-- [Scincidae](scincidae.html) [incl. Feyliniidae] (skinks; skinkit) [124/1206] | --o Cordyloidea [Cordyliformes Fitzinger, 1826] | |-- Cordylidae Mertens, 1937 (non Gray, 1838?, non Fitzinger, 1826?) (Girdled Lizards, vyöliskot) | -- [Gerrhosauridae](gerrhosauridae.html) (kilpiliskot) --o Lacertoidea Camp, 1923 (sisiliskonsukuiset liskot) |--o Xantusiomorpha | |?-o †Hodzhakuliidae Alifanov, 1993 sensu Alifanov, 2016 [previously treated as Anguimorpha incertae sedis] | | |-- †Bagaluus primigenius Alifanov, 2016 | | --+-- †_Hodzhakulia magna_ Nessov, 1985 | | --+-- †“Platynotoides” altidentatus Alifanov, 2016 [Platynotoides Alifanov, 2016 (preocc.), non Platynotoides Kaszab, 1975 (a beetle)] | | --+-- †_Carnoscincus eublepharus_ Alifanov, 2016 | | --o †Pachygenys Gao & Cheng, 1999; E.Cret. EAs. | | |-- †P. thlastesa Gao & Cheng, 1999 | | -- †_P. adachii_ Ikeda, Ota & Saegusa, 2015 | -- Xantusiidae (yöliskot) --o Lacertiformes Estes, de Queiroz & Gauthier, 1988 (sisiliskomaiset liskot) |--+?-o †Eolacertidae Cernansk� & Smith, _in press_ (2017) | | |-- †_Eolacerta robusta_ Nöth, 1940 [however, see [Anguimorpha](../anguimorpha/anguimorpha.html) for alternative position] | | -- †Stefanikia siderea Cernansk� & Smith, in press (2017) | -- [Lacertidae](lacertidae%5F1.html) Bonaparte, 1831 (sisiliskot) [26/273] --+--o †Meyasaurus Vidal, 1915 [Ilerdaesaurus] | |-- †M. faurai Vidal, 1915 (emend. von Huene, 1952) | |-- †M. diazromeri Evans & Barbadillo, 1997 | |-- †M. crusafonti (Hoffstetter, 1966) | -- †_M. unensis_ (Richter, 1994) --+?- †Dalinghosaurus longidigitus Ji, 1998 --+--o Teiioidea Estes, de Queiroz & Gauthier, 1988 (_Teju_-like lisards; teijunsukuiset liskot) | |?- †_Haptosphenus placodon_ Estes, 1964 | --+?- †Adamisaurus magnidentatus Sulimski, 1972 [Adamisauridae Sulimski, 1978] | |-- Teiidae Gray, 1827 (tejus; teijut) [Recent: 9 genera / 116 species] | -- [Gymnophthalmidae](gymnophthalmidae/gymnophthalmidae.html) --o †Boreoteiioidea Nydam, Eaton & Sankey, 2007 |?- †Tijubina pontei Bonfim-J�nior & Marques, 1997 |?- †Olindalacerta brasiliensis Evans & Yabumoto, 1998 |-- †Kuwajimalla kagaensis Evans & Manabe, 2008 |-- †Tianyusaurus zhengi Lü et al., 2008 |-- †Mongolochamopidae Alifanov, 1993 |-- †Macrocephalosauridae Sulimski, 1975 `-- †Polyglyphanodontidae Gilmore, 1943b

| After Alifanov, 2016 **<==**o Scincomorpha |--o Leptoglossa | | --+-- †Globauridae | | `--+-- †Ardeosauridae | | `-- †Carusiidae | `--+--o ?Cordyloidea | | | -- †Paramacellodidae | | `-- Cordylidae | `--o ?Scincoidea | | -- Acontiidae | `-- Scincidae `--o Xantusiomorpha | -- Xantusiidae `--+-- Dibamidae `--+--o †Eoxantidae | `-- †Eoxanta `--+--o †Slavoiidae | `-- †Slavoia `--o †Hodzhakuliidae | -- †Bagaluus_ `--+-- †_Hodzhakulia `--+-- †“Platynotoides” `--+-- †Carnoscincus `-- †Pachygenys_ Reference(s): Alifanov, V. R., 2016: Lizards of the family Hodzhakuliidae (Scincomorpha) from the lower Cretaceous of Mongolia.–Paleontological Journal: Vol. 50, #5, pp. (504-513) [**doi: 10.1134/s0031030116050038**] | After Kearney, 2003 **<==**o Scincomorpha |--o Cordyliformes | `--o Cordyloidea | | -- Cordylidae | `-- Gerrhosauridae `--+--o Scincoidea | | ?- †_Dimekodontosaurus | | ?- †Hymenosaurus | | ?- †Parmeosaurus | `-- Scincidae `--+--o †Eichstaettisauridae | | -- †Eichstaettisaurus | `-- †Globaura_ `--+-- †_Eoxanta `--+-- Xantusiidae | --o Lacertiformes | | -- Lacertidae Bonaparte, 1831 | `--+-- †Meyasaurus | `-- Teiidae `--+--o †Slavoiidae | `-- †Slavoia_ | --o †Adamisauridae | `-- †_Adamisaurus `--+-- †Polyglyphanodontidae `--+-- †Sineoamphisbaena `-- †Macrocephalosauridae Reference(s): Kearney, M., 2003: The phylogenetic position of Sineoamphisbaena hexatabularis reexamined.–Journal of Vertebrate Paleontology: Vol. 23, #2, pp. 394-403 | | --------------------------------------------------------------- | | ------------------ | | ---------------------- | | ---------------- | --------------------- | | | -------------------- | | ---------------------------------- | ----------------- | | ------------- | -------------------------------------------------------- | ------------------------------------------------------------------------------------------------------------- | ------------------------------------------------ | ---------------------------------------------------- | ----------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------- | ---------------------------------------------------------------- | ----------------- | | -------------------------------- | ----------------------------------------------------------------- | | ----------------------- | | ------------------ | | ------------------ | -------------------------------------------------------------------- | | ----------------------- | ------------------------------------------------------------------------------------------------ | ----------------- | | ---------------------------------------------------- | --------------------- | ---------------------------------------------------------------------------- | --------------- | ------------------ | --------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------- |

Reference(s):

• Alifanov, V. R., 2000: The fossil record of Cretaceous lizards from Mongolia. 368-389
  in Benton, M. J., Shishkin, M. A., Unvin, D. M. & Kurochkin, E. N., (eds.) 2000: The Age of Dinosaurs in Russia and Mongolia.
  –Cambridge University Press, Cambridge, 2000, xxxix-696
• Alifanov, V. R., 2016: Lizards of the family Hodzhakuliidae (Scincomorpha) from the lower Cretaceous of Mongolia.
  –Paleontological Journal: Vol. 50, #5, pp. 504-513 [**doi: 10.1134/s0031030116050038**]
• Carroll, R. L., 1988: Vertebrate paleontology and evolution.
  –W. H. Freeman and company, New York, 1988, 698.
• Carroll, R. L., 1988: Appendix. 594-648.
  in Carroll, R. L. 1988: Vertebrate paleontology and evolution.
  –W. H. Freeman and company, New York, 1988, 698.
• Cernansk�, A. & Smith, K. T., 2017: Eolacertidae: a new extinct clade of lizards from the Palaeogene; with comments on the origin of the dominant European reptile group � Lacertidae.
  –Historical Biology: Vol. 30, #7, pp. 994-1014 [**doi: 10.1080/08912963.2017.1327530**]
• Evans, S. E. & Barbadillo, L. J., 1997: Early Cretaceous lizards from Las Hoyas, Spain.
  –Zoological Journal of the Linnean Society: Vol. 119, #1, pp. 23-49
• Evans, S. E. & Chure, D. C., 1998: Paramacellodid lizard skulls from the Jurassic Morrison Formation at Dinosaur National Monument, Utah.
  –Journal of Vertebrate Paleontology: Vol. 18, #1, pp. 99-114
• Evans, S. E. & Manabe, M., 2008: An early herbivorous lizard from the lower Cretaceous of Japan.
  –Palaeontology: Vol. 51, #2, pp. 487-498 [doi: 10.1111/j.1475-4983.2008.00759.x]
• Folie, A. & Codrea, V., 2005: New lissamphibians and squamates from the Maastrichian of Hateg Basin, Romania.
  –Acta Palaeontologica Polonica: Vol. 50, #1, pp. 57-71
• Folie, A., Sig�, B. & Smith, T., 2005: A new scincomorph lizard from the Palaeocene of Belgium and the origin of Scincoidea in Europe.
  –Naturwissenschaften: Vol. 92, #11, pp. 542-546 [doi: 10.1007/s00114-005-0043-4]
• Gao, K.-Q. & Cheng, Z., 1999: A new lizard from the Lower Cretaceous of Shandong, China. –Journal of Vertebrate Paleontology: Vol. 19, #3, pp. 456-465
• Gao, K.-Q. & Norell, M. A., 2000: Taxonomic composition and systematics of Late Cretaceous lizard assemblages from Ukhaa Tolgod and adjacent localities, Mongolian Gobi Desert.
  –Bulletin of the American Museum of Natural History: #249, pp. 1-118
• Ikeda, T., Ota, H. & Saegusa, H., 2015: A new fossil lizard from the Lower Cretaceous Sasayama Group of Hyogo Prefecture, western Honshu, Japan.
  –Journal of Vertebrate Paleontology: Vol. 35, #1, pp. e885032 (1-5) [**doi: 10.1080/02724634.2014.885032**]
• Kearney, M., 2003: The phylogenetic position of Sineoamphisbaena hexatabularis reexamined.
  –Journal of Vertebrate Paleontology: Vol. 23, #2, pp. 394-403
• Nydam, R. L., 2002: Lizards of the Mussentuchit Local Fauna (Albian-Cenomanian Boundary) and comments on the evolution of the Cretaceous lizard fauna of North America.
  –Journal of Vertebrate Paleontology: Vol. 22, #3, pp. 645-660
• Nydam, R. L. & Cifelli, R. L., 2002: Lizards from the Lower Cretaceous (Aptian-Albian) Antlers and Cloverly Formations.
  –Journal of Vertebrate Paleontology: Vol. 22, #2, pp. 286-298
• Reynoso, V.-H. & Callison, G., 2000: A new scincomorph lizard from the Early Cretaceous of Puebla, Mexico.
  –Zoological Journal of the Linnean Society: vol. 130, #2, pp. 183-212.
• Rieppel, O. 1994: The Lepidosauromorpha: an overview with special emphasis on the Squamata. 23-37.
  in Fraser, N. C. & Sues, H-D. 1994: In the Shadow of the Dinosaurs.
  –Cambridge University Press, New York. 1994
• Sim�es, T. R., Caldwell, M. W. & Kellner, A. W. A., 2014: A new Early Cretaceous lizard species from Brazil, and the phylogenetic position of the oldest known South American squamates.
  –Journal of Systematic Palaeontology: Vol. 13, #7, pp. 601-614 [**doi: 10.1080/14772019.2014.947342**]
• Uetz, P., Etzold, T. & Chenna, R., 1995-2000: The Reptile Database
  –iNet: http://www.reptile-database.org/