Anapsida (original) (raw)

After Tsuji & Müller, 2009

**<==o †Parareptilia Olson, 1947 sensu Tsuji & Müller, 2009 [Anapsida Osborn, 1903 [paraphyletic]; Proganosauria; ] (anapsids, parareptiles; ohimoaukottomat amniootit, parareptiilit) |--o †Mesosauridae Baur, 1889 | |-- †Brazilosaurus sanpauloensis | |-- †Stereosternum tumidum Cope, 1886 | --o †_Mesosaurus_ [_Ditrichosaurus_, _Noteosaurus africanus_ Broom, 1913, _Notosaurus_] | |-- †_M. tenuidens_ Gervais, 1865 | -- †M. brasiliensis [jr. synon. to M. tenuidens??] --o Parareptilia Olson, 1947 _sensu_ deBraga & Reisz, 1996 |--+-- †_Eunotosaurus africanus_ Seeley, 1892 _sensu_ Modesto, 2000 | ?- Testudinata Klein, in Behn, 1760 sensu Lyson, Bever, Bhullar, Joyce & Gauthier, 2010, and Lyson, Bever, Scheyer, Hsiang & Gauthier, 2013 (turtles & tortoises) |--o †Millerettidae Watson, 1957 [Millerosauria, Millerosauroidea] | |?- †Heleophilus Broom, 1909 | |?- †Milleretoides acutirostris Broom, 1941 | |?- †Nanomilleretta kitchingi Broom, 1948 | |-- †Milleretta rubidgei Broom, 1938 [Millerina Broom, 1938; Millerettoides platyceps Broom, 1948; Millerettops kitchingi Broom, 1948] | --+-- †_Broomia perplexa_ Watson, 1914 | --+-- †Millerosaurus ornatus Broom, 1948 | --+-- †_Milleropsis pricei_ Gow, 1972 [=? _Millerosaurus pricei_] | -- †Millerosaurus nuffieldi Watson, 1957 --o †Ankyramorpha deBraga & Reitz, 1996 _sensu_ Tsuji & Müller, 2009 |--o †Lanthanosuchoidea Efremov, 1946 _sensu_ Debraga & Reisz, 1996 | ==**o †“Lanthanosuchidae” Efremov, 1946 [paraphyletic ( Tsuji & Müller, 2009)] | |-- †Lanthaniscus efremovi Ivakhnenko, 1980 | |-- †Lanthanosuchus watsoni Efremov, 1946 | |-- †Chalcosaurus rossicus v. Meyer, 1866 | --o †Acleistorhinidae Daly, 1969 | |-- †_Colobomycter pholeter_ Vaughn, 1958 | -- †Acleistorhinus pteronicus Daly, 1969 --+--o †Nyctiphruretia Efremov, 1938 _sensu_ Lee, 1993 [paraphyletic?] | |?- †_Tokosaurus perforatus_ Tverdochlebova & Ivachnenko, 1984 | --o †Nyctiphruretidae Efremov, 1938 | |?-†Nyctiboetus liteus Chudinov 1955 [a nyctiphruretid according to Paleobiology Database] | |-- †Abyssomedon williamsi MacDougall & Reisz, 2014 | --o †_Nyctiphruretus_ Efremov, 1938 | |-- †_N. acudens_ Efremov, 1938 [_N. acrudens_ (lapsus calami)] | |-- †_N. ineptus_ (=? _Nycteroleter ineptus_ Efremov, 1938) | -- †N. optabilis Bulatov, 2002 |--o †Bolosauria Kuhn, 1959 sensu Modesto, Scott, MacDougall, Sues, Evans & Reisz, 2015 | |-- †Erpetonyx arsenaultorum Modesto, Scott, MacDougall, Sues, Evans & Reisz, 2015 | --o †Bolosauridae Cope, 1878 _sensu_ Berman et al., 2000 | |?- †**Bolosauridae** _genera_ et _species_ indet. [China] | |--o †_Bolosaurus_ Cope, 1878; L. Perm. NA. | | |?- †_B. major_ | | |?- †_B. rapidens_ | | |-- †_B. striatus_ Cope, 1878 | | -- †B. grandis Reisz, Barkas & Scott, 2002 | |-- †Eudibamus cursoris Berman et al., 2000, L. Perm. CEu. | --o †_Belebay_ U. Perm. EEu. | |-- †_B. vegrandis_ | -- †B. maximi `-- †Procolophonomorpha Romer, 1964 sensu Lee, 1995

| After Modesto, Scott, MacDougall, Sues, Evans & Reisz, 2015 **<==**o Parareptilia |-- †Mesosauridae `--+--o †Millerosauria | | -- †Eunotosaurus_ | `-- †Millerettidae `--+--o †Bolosauria | | -- †Erpetonyx | `-- †Bolosauridae `--+-- †Australothyris `--+-- †Lanthanosuchia `--+-- †Microleter `--+-- †Nyctiphruretidae `--+-- †Procolophonidae `-- †Pareiasauromorpha | After Lyson, Bever, Bhullar, Joyce & Gauthier, 2010 **<==**o Anapsida |-- †Mesosauridae `--o Parareptilia | -- †Millerettidae `--+--o †Lanthanosuchoidea | | -- †_Acleistorhinus | `-- †Lanthanosuchus_ `--+--o Pan-Testudines | | -- †_Eunotosaurus | `-- Testudinata `--+-- †Nyctiphruretus `--+--+-- †Bolosauridae | `-- †Procolophonoidea `--+-- †Nycteroleteridae `-- †Pareiasauria | After Berman et al., 2000, Lee, 1993, 1996,, and Modesto 1999, 2000 **<==**o Anapsida |-- †Mesosauridae `--o Parareptilia | -- †Millerettidae `--+-- †Bolosauridae `--+-- †Eunotosaurus_ sensu Modesto, 2000 `--o Procolophonomorpha | ?- †_Australothyris `--+-- †Procolophonoidea `--+-- †Nycteroleteridae `-- †Pareiasauria | | ---------------------------------------------------------------------------------------------------------------------------------- | | ------------------ | ---------------------------------------------- | | --------------- | ----------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------- | ----------------------------------------------------------------------------------------------------------------- | --------------------------------------------------- | | -------------------- | ------------------------------------------------------- | | ------------------ | ----------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------- | --------------------------------------------------------------------------------------------------------------------------------------------------------- | --------------------------------------------------------------------------------------------------------------------------------- | -------------------------------------------------------------------------------------------------------------------------------------------------------------------- | ------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------ |

Reference(s):

• Berman, D. S., Reisz, R. R., Scott, D., Henrici, A. C., Sumida, S. S. & Martens, T. 2000: Early Permian bipedal reptile.
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• Carroll, R. L., 1988: Vertebrate paleontology and evolution.
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• Carroll, R. L., 1988: Appendix. 594-648.
  in Carroll, R. L. 1988: Vertebrate paleontology and evolution.
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