†Procolophonoidea (original) (raw)

After Spencer & Benton, 2000, Spencer & Sues, 2000, Reisz & Scott, 2002, de Beaga, 2003, Modesto, Damiani, Neveling & Yates, 2003, and Cisneros, Damiani, Schultz, de Rosa, Schwanke, Neto & Aurélio, 2004

**<==**o †Procolophonoidea Romer, 1956 |?- †Xenodiphyodon petraios Sues & Olsen, 1993 |--o Owenettidae Broom, 1939 | |-- †Barasaurus besairei Piveteau, 1955 | |-- †Ruhuhuaria reiszi Tsuji, Sobral & M�ller, 2013 | --+-- †_Owenetta rubidgei_ Broom, 1939 [_Colubrifer campi_ Carroll, 1982] | --+-- †Saurodektes rogersorum (Modesto, Damiani, Neveling & Yates, 2003) Modesto, Damiani, Neveling & Yates, 2004 [Saurodectes rogersorum_ Modesto, Damiani, Neveling & Yates, 2003; non †Saurodectes Rasnitsyn & Zherikhin, 2000, a extinct louse (Insecta: Phthiriaptera)] | --+-- †_Candelaria barbouri_ Price, 1946 | -- †“Owenetta” kitchingorum Reisz & Scott, 2002 --o †Procolophonidae Cope, 1889 _sensu_ S�il�, 2009 [_non_ Seeley, 1888; _non_ Lydekker, 1890] [incl. †Sclerosauridae] |-- †_Coletta seca_ Gow, 2000 |?- †_Kinelia broomi_ Bulatov, 2002 |-- †_Pintosaurus magnidentis_ Piñeiro, Rojas & Ubilla, 2004 --+-- †_Sauropareion anoplus Modesto, Sues & Damiani, 2001 --+-- †_Gomphiosauridion baileyae_ Sues & Olsen, 1993 |--o †Theledectinae Cisneros, 2008 | |-- †_Theledectes perforatus_ (Gow, 1977) Modesto & Damiani, 2003 [_Thelegnathus_, in partim] | --o †Eumetabolodon Li, 1983 [Eumetabolodan Cisneros, 2008 (lapsus calami)] | |?- †E. bathycephalus Li, 1983 | -- †_E. dongshengensis_ Li, 1983 |?-o Spondylolestinae Ivakhnenko, 1979 [paraphyletic? assemblage of basal procolophonoids] | |-- †_Neoprocolophon asiaticus_ Young, 1957 | |-- †_Phaanthosaurus ignatjevi_ Tchudinov & Vjuschkov, 1956 | |-- †_Procolophonoides baini_ (Broom, 1905) Ivakhnenko, 1982 [_Procolophon baini_ Broom, 1905; _Procolophon trigoniceps_ Owen; Colbert & Kitching, 1975] | ?- †Spondylolestes rubidgei Broom, 1937 --+--o †_Contritosaurus_ Ivakhnenko, 1974 [paraphyletic **Spondylolestinae**] | |-- †_C. convector_ Ivakhnenko, 1974 | -- †C. simus Ivakhnenko, 1974 --+--o †_Tichvinskia_ Tchudinov & Vjuschkov, 1956 | |-- †_T. vjatkensis_ Tchudinov & Vjuschkov, 1956 | -- †T. jugensis Tchudinov & Vjuschkov, 1956 --+--+-- †_Koiloskiosaurus coburgensis_ v. Huene, 1913 [_Koiliskiosaurus_; _Koiloskiosaurus koburgensis_ (lapsus calami?)] | --+?-o †“Sclerosauridae” [treated as pareiasauroids by Lee (1995)] | | |?- †Basileosaurus freyi Wiedersheim, 1879 | | -- †_Sclerosaurus armatus_ v. Meyer, 1859 [_Aristodesmus_] | --+?- †Thelegnathus browni Broom, 1905 (nomen dubium sensu Modesto & Damiani, 2003) | |?- †Thelerpeton oppressus (Gow, 1977) Modesto & Damiani, 2003 [Thelegnathus, in partim] | |?- †Thelephon contritus (Gow, 1977) Modesto & Damiani, 2003 [Thelegnathus, in partim] | |?- †Teratophon spinigenis (Gow, 1977) Modesto & Damiani, 2003 [Thelegnathus, in partim] | --o †Leptopleuroninae Ivakhnenko, 1979 | |?- †_Paoteodon huanghoensis_ Chow & Sin, 1960 (_nomen dubium_?) | |?- †_Scoloparia glyphanodon_ Sues & Baird, 1998 | |-- †_Libognathus sheddi_ Small, 1997 | --o †Leptopleuronini | |-- †Hypsognathus fenneri Gilmore, 1928 | -- †_Leptopleuron lacertinum_ Owen, 1851 [_Telerpeton elginense_ Mantell, 1852] --o †Procolophoninae Lydekker, 1890 (non Seeley, 1888 emend. Romer, 1956) Ivakhnenko, 1979 |--o †Anomoiodon von Huene, 1939 | |-- †A. liliensterni von Huene, 1939 | |-- †A. huenei | -- †_A. krejcii_ |-- †_Burtensia burtensis_ (Otschev, 1958) Ivakhnenko 1975 |-- †_Insulophon morachovskayae_ Novikov, 1992 [jr. synon. of _Orenburgia bruma_?] |--o †_Kapes_ Ivakhnenko, 1975 [?_Macrophon_, ?_Orenburgia_ Ivakhnenko, 1975, _Samaria_ Novikov, 1991, _Lestanshoria_ Novikov, 1991] | |-- †_K. amaenus_ Ivakhnenko, 1975 | |-- †_K. majmesculae_ (Otschev, 1968) [_Tichvinskia majmesculae_ Otschev, 1968, _Orenburgia majmesculae_ (Otschev, 1968)] | -- †K. serotinus Novikov, 1991 |-- †Lestanshoria massiva Novikov, 1991 [jr. synon. of Orenburgia?] |-- †Macrophon komiensis Ivakhnenko, 1975 |-- †Microthelodon parvus (Broom, 1936) Romer, 1966 [Thelegnathus parvus Broom, 1905] |-- †Myocephalus crassidens Broom, 1936 |--o †Orenburgia Ivakhnenko, 1975 | |-- †O. enigmatica (Tchudinov & Vjuschkov, 1956) [Tichvinskia enigmatica Tchudinov & Vjuschkov, 1956] | -- †_O. bruma_ Ivakhnenko, 1983 |--o †_Procolophon_ Owen, 1876 | |-- †_P. trigoniceps_ Owen, 1876 [_P. minor_ Owen, 1876; _P. griersoni_ Seeley, 1878; _P. cuneiceps_ Seeley, 1878; _P. laticeps_ Seeley, 1878; _P. platyrhinus_ Seeley, 1905; _P. sphenorhinus_ Seeley, 1905] | |-- †_P. pricei_ Lavina, 1983 | -- †P. brasiliensis Cisneros & Schultz, 2002 |-- †Samaria concinna (Ivakhnenko, 1975) [Orenburgia concinna Ivakhnenko, 1975, Samaria Novikov, 1991] |-- †Soturnia caliodon Cisneros & Schultz, 2003 `-- †Timanophon raridentatus Novikov, 1991

Procolophonoidea incertae sedis:

†_Acadiella psalidodon_ Sues & Baird, 1998
†_Colognathus obsurus_ Case, 1928
†_Estheriophagus chatangensis_ Novojilov, 1948
†_Haligonia bolodon_ Sues & Baird, 1998
†_Myognathus_
†_Santaisaurus yuani_ Koh, 1940
†_Suchonosaurus minimus_ Tverdokhlebova & Ivakhnenko, 1994 [a non-procolophonoid? (Cisneros, 2008)]
†_Vitalia grata_ Ivakhnenko, 1973 [a non-procolophonoid?]

After Cisneros, 2008 (fig. 4)

**<==**o †Procolophonidae |-- †Coletta seca |-- †Pintosaurus magnidentis |-- †Sauropareion anoplus --+-- †_Phaanthosaurus_ spp. --+--o †Theledectinae | |-- †Theledectes perforatus | -- †“_Eumetabolodon_” _dongshengensis_ --+-- †Tichvinskia vjatkensis --+--o †Procolophoninae | |--+-- †_Timanophon raridentatus_ | | --+-- †Kapes spp. | | -- †_Thelephon contritus_ | --+-- †Eumetabolodon bathycephalus | --+-- †_Procolophon trigoniceps_ | |-- †_Thelerpeton oppressus_ | -- †Teratophon spinigenis --o †Leptopleuroninae |-- †_Pentaedrusaurus ordosianus_ --+-- †Neoprocolophon asiaticus --+--+-- †_Sclerosaurus armatus_ | -- †Scoloparia glyphanodon --+-- †_Leptopleuron lacertinum_ --+-- †Hypsognathus fenneri `-- †Soturnia caliodon

Reference(s):

Reference(s):