†Paracryptodira (original) (raw)

After Meylan & Ganko, 1997, Lyson & Joyce, 2010, and Lyson & Joyce, 2011

**<==o †Paracryptodira [=? Pleurosternoidea Hay, 1930] |?-o †Solemydidae de Lapparent de Broin & Murelaga, 1996 | |?- †Plastremys lata Owen, in Parkinson, 1881 [Hylaeochelys lata (Owen) Milner, 2004] | |--o †Helochelydra Nopcsa, 1928 [=? †Helochelys] | | |-- †H. nopcsai de Lapparent de Broin & Murelaga, 1999 | | |-- †H. bakewelli (Mantell, 1833) Milner, 2004 | | -- †_H. anglica_ (Lydekker, 1889b) Milner, 2004 | |-- †_Naomichelys speciosa_ Hay, 1908 | -- †Solemys vermiculata de Lapparent de Broin and Murelaga 1996 |--o †Pleurosternidae Cope, 1868 [incl. ?Chengyuchelyidae Yeh, 1990] | |?-o †Chengyuchelys Young & Chow, 1953 [Chengyuchelyidae Yeh, 1990] | | |-- †C. baenoides Young & Chow, 1953 | | -- †_C. zigongensis_ Yeh 1982 | |-- †_Dorsetochelys delairi_ Evans & Kemp, 1976 [**Cryptodira** _incertae sedis_ (Milner, 2004)] | |?- †_Uluops uluops_ Bakker, 1990 | |--o †_Pleurosternon_ Owen, 1853 [**[taxonomic history](history%5FPleurosternon.html)**] | | |-- †_P. bullockii_ (Owen, 1842) Lydekker & Boulenger, 1887 [**[taxonomic history](history%5FPleurosternon.html#P%5Fbullockii)**] | | -- †P. portlandicum Lydekker, 1889 | |?-o †Compsemys Leidy, 1856; E.Cret.-Pal. NAm. Eu. | | |-- †C. bakewelli (Mantell, 1833) [Helochelydra bakewelli; Tretosternum bakewelli; Trionyx bakewelli] | | -- †_C. victa_ Leidy, 1856 [_Compsemys obscura_ Leidy, 1856; _Compsemys parva_ Hay, 1910; _Compsemys puercensis_ Gilmore, 1919; _Compsemys torrejonensis_ Gilmore, 1919; _Compsemys vafer_ Hay, 1910; _Emys obscurus_ Leidy, 1856; _Glyptops depressus_ Hay, 1908] | --+--o †“Glyptopsinae” Marsh, 1890 [Glyptopsidae Marsh, 1890] | | |?- †Mesochelys durlstonensis Evans & Kemp, 1975 [treated as a synonym of Pleurosternon by Milner (2004)] | | --o †_Glyptops_ Marsh, 1890 [**[taxonomic history](history%5FGlyptops.html)**] | | |?- †“_Glyptops_” _typocardium_ (Seeley, 1869) Milner, 2004 [**[taxonomic history](history%5FGlyptops.html#G%5Ftypocardium)**] | | |-- †_G. plicatulus_ (Cope, 1877) Marsh, 1890 [**[taxonomic history](history%5FGlyptops.html#G%5Fplicatulus)**] | | -- †G. ruetimeyeri (Lydekker, 1889b) [**taxonomic history] | --o †“Dinochelinae” | |-- †_Desmemys bertelsmanni_ Wegner, 1911 | -- †Dinochelys whitei Gaffney, 1979 --o †Baenoidea [Baenidae _sensu lato_] [paraphyletic??] |?- †[Macrobaenidae](macrobaenidae.html) Sukhanov, 1964 --o †Baenidae Cope, 1882 sensu stricto |?- †Goleremys mckennai Hutchison, 2005 |-- †Arundelemys dardeni Lipka, Therrien, Weishampel, Jamniczky, Joyce, Colbert & Brinkman, 2006 --+-- †_Trinitichelys hiatti_ Gaffney, 1972 --+--o †Neurankylus Lambe, 1902 [Charitemys Hay, 1908] [Neurankylidae] | |-- †N. eximius Lambe, 1902 [Baena fluviatilis Parks 1933; Charitemys captans Hay, 1908; Neurankylus baueri Gilmore, 1916; Thescelus fluviatilis Parks, 1933] | -- †_N. wyomingensis_ Gilmore, 1919 --+-- †Hayemys latifrons Gaffney, 1972 |-- †Thescelus insiliens Hay, 1908 [Baena longicauda Russell, 1934; Thescelus hemispherica Gilmore, 1935; Thescelus rapiens Hay, 1908] |-- †“Denazinemys_” ornata (Gilmore, 1935) Lucas & Sullivan, 2006 [Baena ornata Gilmore, 1935] --o †Baenodda Gaffney & Meylan, 1988 [Eubaeninae] |?- †“_Baena_” _hatcheri_ Hay, 1901 |?- †“_Baena_” _marshi_ Hay, 1904 |--+-- †“_Baena_” _hayi_ Gilmore, 1916 | --+--o †_Plesiobaena Gaffney, 1972 | | |-- †P. antiqua (Lambe, 1902) [Baena antiqua Lambe, 1902; Baena antiqua Lambe; Russell, 1934; Plesiobaena antigua (Lambe); Gaffney, 1972 (in partim)]; U. Cret. NA. | | --+-- †_P. sp._ U. Cret. NA. [Maastrichian species (Brinkman, 2003)] | | -- †P. putorius Gaffney, 1972; Pal. NA. [=? Cedrobaena putorius] | --+-- †_Peckemys brickman_ | --+-- †Cedrobaena putorius [=? Plesiobaena putorius Gaffney, 1972] | --+-- †_Gamerabaena sonsalla_ Lyson & Joyce, 2010 | --o †Palatobaena | |-- †P. cohen Lyson & Joyce, 2009 | --+-- †_P. gaffneyi_ | -- †P. bairdi --+--+-- †_Stygiochelys estesi_ Gaffney & Hiatt, 1971 | --+-- †Baena arenosa Leidy, 1890 | --o †_Chisternon_ Leidy, 1872 | |-- †_C. undatum_ (Leidy, 1871) | -- †C. hebraicum Cope 1872 --+-- †_Eubaena cephalica_ (Hay, 1904) Hay, 1908 [_Baena cephalica_ Hay, 1904] --+-- †Denazinemys nodosa (Gilmore, 1916) Lucas & Sullivan, 2006 [Baena nodosa Gilmore, 1916] --o †_Boremys_ Lambe, 1906 |-- †_B. pulchra_ Lambe, 1906 -- †B. grandis Gilmore, 1935

| After Lyson & Joyce, 2011 <==o †Paracryptodira |-- †Dinochelys whitei `--+-- †Compsemys victa_** **`--+-- †_Uluops uluops |--o †Pleurosternidae | | -- †Dorsetochelys delairi | `--+-- †Pleurosternon_** **| `-- †_Glyptops `--o †Baenidae** **|-- †Arundelemys dardeni_** **`--+-- †_Trinitichelys hiatti `--+-- †Neurankylus eximius_** **`--+-- †_Hayemys latifrons |-- †“_Denazinemys_” ornata |--+--+-- †Stygiochelys estesi | | `--+-- †Baena arenosa_** **| | `-- †_Chisternon undatum | `--+-- †Eubaena cephalica_** **| `--+-- †_Denazinemys nodosa | `--o †Boremys_** | | -- †B. pulchra **| `-- †_B. grandis `--+-- †Plesiobaena antiqua_** **`--+-- †_Peckemys brickmani `--+-- †Cedrobaena putorius_** **`--+-- †_Gamerabaena sonsalla `--o †Palatobaena_** |-- †P. cohen **`--+-- †_P. gaffneyi `-- †_P. bairdi_** **Reference(s): Lyson, T. R. & Joyce, W. G., 2011: Cranial Anatomy and Phylogenetic Placement of the Enigmatic Turtle _Compsemys victa_ Leidy, 1856.–Journal of Paleontology: Vol. 85, #4, pp. 789-801 [doi: 10.1666/10-081.1]** | After Lyson & Joyce, 2010 **<==o †Paracryptodira** **|-- †_Pleurosternon_** **|-- †_Glyptops_** **`--o †Baenidae |--+-- †Trinitichelys hiatti | `-- †Neurankylus eximius_** **`--+-- †_Thescelus insiliens |-- †Hayemys latifrons `--o †Baenodda** **|--+-- †“Baena_” hayi** **| `--+-- †_Plesiobaena antiqua | `--+-- †Gamerabaena sonsalla_** **| `--o †_Palatobaena | | -- †P. cohen | `--+-- †P. gaffneyi_** **| `-- †_P. bairdi `--+--+-- †_Boremys pulchra_** **| | -- †_Boremys grandis_** **| | -- †_Eubaena cephalica_** **| `-- †“_Baena_” hatcheri `--+-- †Stygiochelys estesi_** **`--+-- †_Baena arenosa `-- †_Chisternon undatum_** **Reference(s): Lyson, T. R. & Joyce, W. G., 2010: A new baenid turtle from the upper Cretaceous (Maastrichtian) Hell Creek Formation of North Dakota and a preliminary taxonomic review of Cretaceous Baenidae.–Journal of Vertebrate Paleontology: Vol. 30, #2, pp. (394-402) [doi: 10.1080/02724631003618389]** | After Lipka, Therrien, Weishampel, Jamniczky, Joyce, Colbert & Brinkman, 2006 **<==o †Paracryptodira** **|-- †_Arundelemys dardeni_** **`--+--o †Pleurosternidae | | -- †Pleurosternon | `--+-- †Glyptops_** **| `-- †_Dinochelys whitei `--o †Baenidae** **|-- †Neurankylus eximius_** **`--+-- †_Trinitichelys hiatti `--+-- †Plesiobaena antiqua_** **`--+-- †_Boremys pulchra `--+-- †Baena arenosa_** **`-- †_Chisternon undatum Reference(s): Lipka, T. R., Therrien, F., Weishampel, D. B., Jamniczky, H. A., Joyce, W. G., Colbert, M. W. & Brinkman, D. B., 2006: A new turtle from the Arundel clay facies (Potomac Formation, Early Cretaceous) of Maryland, U.S.A.–Journal of Vertebrate Paleontology: Vol. 26, #2, pp. 300-307 [doi: 10.1671/0272-4634(2006)26[300:antfta]2.0.co;2] | | ------------------------------------------------------------------------------------------------------------------------------------------------------------------------------- | ------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------ | ------------------------------- | ---------------------------------------------------------------------------------------------------------------------------------------------- | --------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------- | ---------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------- | -------------------------------------------------------------------------------------------------------------------------------------- | ---------------------------- | ---------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------- | --------------------------------------------------------------------------------------------------------------------------------------------------------------------------- | ------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------------ |

Reference(s):

• Brinkman, D. B., 2003: Anatomy and systematics of Plesiobaena antiqua (Testudines: Baenidae) from the Mid-Campanian Judith River Group of Alberta, Canada.
  –Journal of Vertebrate Paleontology: Vol. 23, #1, pp. 146-155
• Brickman, D. B., & Nicholls, E. L., 1993: The skull of Neurankylus eximius (Testudines: Baenidae) and a reinterpretation of the relationships of this taxon.
  –Journal of Vertebrate Paleontology: Vol. 13, #3, pp. 273-281
• Brickman, D. B., Stadtman, K. & Smith, D., 2000: New material of Dinochelys whitei Gaffney, 1979, from the Dry Mesa Quarry (Morrison Formation, Jurassic) of Colorado.
  –Journal of Vertebrate Paleontology: Vol. 20, #2, pp. 269-274
• Lipka, T. R., Therrien, F., Weishampel, D. B., Jamniczky, H. A., Joyce, W. G., Colbert, M. W. & Brinkman, D. B., 2006: A new turtle from the Arundel clay facies (Potomac Formation, Early Cretaceous) of Maryland, U.S.A.
  –Journal of Vertebrate Paleontology: Vol. 26, #2, pp. 300-307 [**doi: 10.1671/0272-4634(2006)26[300:antfta]2.0.co;2**]
• Lyson, T. R. & Joyce, W. G., 2010: A new baenid turtle from the upper Cretaceous (Maastrichtian) Hell Creek Formation of North Dakota and a preliminary taxonomic review of Cretaceous Baenidae.
  –Journal of Vertebrate Paleontology: Vol. 30, #2, pp. (394-402) [**doi: 10.1080/02724631003618389**]
• Lyson, T. R. & Joyce, W. G., 2011: Cranial Anatomy and Phylogenetic Placement of the Enigmatic Turtle Compsemys victa Leidy, 1856.
  –Journal of Paleontology: Vol. 85, #4, pp. 789-801 [**doi: 10.1666/10-081.1**]
• Meylan, P. A. & Ganko, E., 1997: Higher Relationships of Testunides – The extinct and living turtles.
  –iNet: The Tree of Life: [no longer available]
• Milner, A. R., 2004: The turtles of the Purbeck Limestone Group of Dorset, southern England.
  –Palaeontology: Vol. 47, #6, pp. 1441-1467
• Sukhanov, V. R., 2000: Mesozoic turtles of Middle and Central Asia. 309-367
  in Benton, M. J., Shishkin, M. A., Unvin, D. M. & Kurochkin, E. N., (eds.) 2000: The Age of Dinosaurs in Russia and Mongolia.
  –Cambridge University Press, Cambridge, 2000, xxxix-696
• Zonneveld, J.-P., Gunnell, G. F. & Bartels, W. S., 2000: Early Eocene fossil vertebrates from the southwestern Green River basin, Lincoln and Uinta Counties, Wyoming.
  –Journal of Vertebrate Paleontology: Vol. 20, #2, pp. 369-386