John Morse | Clemson University (original) (raw)

Papers by John Morse

Beiträge zur Entomologie, Dec 21, 2023

Based on a recalibrated BEAST diversification time analysis, we provide a revised chronology for ... more Based on a recalibrated BEAST diversification time analysis, we provide a revised chronology for the evolution of major lineages of Trichoptera. Fossil evidence indicates that caddisflies evolved at least by the Norian of Late Triassic (median age 222.6 Ma), compared with our estimate of at least 201.3 Ma. The ancestors of suborders Annulipalpia and Integripalpia also evolved as early as the Norian. Fossil evidence indicates that the ancestor of subterorder Phryganides lived at least by the Aalenian of Middle Jurassic (median age 173.6 Ma), compared with our estimate of at least 174.1 Ma.

Insecta Mundi, Sep 1, 1997

The phylogenetic relationships of the species ofPadunielia are analyzed based on characters oflar... more The phylogenetic relationships of the species ofPadunielia are analyzed based on characters oflarvae, pupae, and adults (mainly male genitalia). The genus is monophyletic and most closely related to Psychomyia, and Metalype in the subfamily Psychornyiinae. Nine species groups are suggested. Eight species, including six new to science, are reported from the People's Republic of China for the first time.

Zoosymposia, Jun 12, 2020

Taxonomy and Basic Description Protoptila morettii was described by John Morse in 1990 from male ... more Taxonomy and Basic Description Protoptila morettii was described by John Morse in 1990 from male and female adults collected in South Carolina. Males measure 3.2 to 3.5 mm (0.13 to 0.14 inches) from front of head to tips of folded wings, with forewings each 2.7 to 3.0 mm (0.11 to 0.12 inches) long. Females measure 3.1 to 4.1mm (0.12 to 0.16 inches) from front of head to tips of folded wings, with forewings each 2.9 to 3.8 mm (0.11 to 0.15 inches) long. The eggs, larvae and pupae of this species remain unknown. Species of Protoptila are members of the subfamily Protoptilinae, family Glossosomatidae, order Trichoptera. The Trichoptera, or caddisflies, are holometabolous insects that are most closely related to Lepidoptera, or moths. Trichoptera eggs, larvae and pupae usually are fully submerged in water and depend on oxygenated water for respiration. Adult caddisflies are aerial/terrestrial, but are rarely found far from water. Adults generally resemble those of their Lepidoptera cousins except, rather than being covered by scales, Trichoptera wings are typically clothed with hair, which is presumably important in repelling water, hence the Latin name trichos (hairy) and ptera (wings). Trichoptera larvae generally resemble those of Lepidoptera except that caddisflies never have fleshy, crochet-bearing prolegs on middle abdominal segments.

Zoosymposia, Jun 12, 2020

Zootaxa, Sep 14, 2017

The Chinese species of the genera Lannapsyche and Marilia (Trichoptera, Odontoceridae) are review... more The Chinese species of the genera Lannapsyche and Marilia (Trichoptera, Odontoceridae) are reviewed, including 1 new species of Lannapsyche (Lannapsyche altilis sp. n.), 1 new Chinese record of Lannapsyche (Lannapsyche chantaramongkolae Malicky 1989, originally described from Thailand), and 3 new species of Marilia (Marilia discaulis, sp. n., Marilia megalopos, sp. n., and Marilia qinlingensis, sp. n.), bringing the Chinese fauna of Lannapsyche to 3 species and of Marilia to 6 species. Previously known species Lannapsyche setschuana Malicky 1995b and Marilia parallela Hwang 1957 are redescribed and reillustrated for both males and females. Updated information about the holotype specimen of Marilia simulans Forsslund 1935 is provided. Distinctive differences of the female genitalia are described, which may add important diagnostic characters for genus and species recognition.

Zenodo (CERN European Organization for Nuclear Research), Dec 31, 2012

Currently, 21 species of the genus Plectrocnemia are known from China. Examination of material co... more Currently, 21 species of the genus Plectrocnemia are known from China. Examination of material collected from 13 provinces of China from 1990-2005 has revealed 9 new species and 3 new records of this genus, bringing the number of Chinese Plectrocnemia species to 33. Newly described species include:

Zoosymposia, Jun 12, 2020

Zootaxa, Aug 1, 2014

Currently, 8 species of the genus Nyctiophylax Brauer are known from China. Examination of materi... more Currently, 8 species of the genus Nyctiophylax Brauer are known from China. Examination of material collected from Guangdong, Guangxi, Jiangxi and Sichuan Provinces during 2004-2005 has revealed 4 new species and 2 new records of this genus, bringing the number of Chinese Nyctiophylax species to 14. Newly described species include: Nyctiophylax (Paranyctiophylax) crinalis n. sp., N. (P.) dactylatus n. sp., N. (P.) orbicularis n. sp., and N. (P.) macrorrhinus n. sp. Nyctiophylax (Paranyctiophylax) sagax Mey and N. (N.) amphonion Malicky & Chantaramongkol are newly recorded for the Chinese fauna, which are re-illustrated and re-described for clear comparisons. The additional collection sites for the previously described species are provided.

Zenodo (CERN European Organization for Nuclear Research), Dec 31, 2010

Six new species of the caddisfly genus Orthotrichia are described and illustrated from China, nam... more Six new species of the caddisfly genus Orthotrichia are described and illustrated from China, namely Or. apophysis Zhou & Yang, sp. nov.; Or. latiramifera Zhou & Yang, sp. nov.; Or. cornuta Zhou &Yang, sp. nov.; Or. discedata Zhou & Morse, sp. nov.; Or. cuspidigera Zhou & Yang, sp. nov.; and Or. subrhomba Zhou & Morse, sp. nov.. One species, Or. terpsichore Malicky & Chantaramongkol, 2007, is recorded for the first time from China, bringing the number of species of the genus known from China to 14. Of these, 10 species are distributed among 3 species groups of Marshall (1979), with 4 species unassigned. The known distributions for all the Chinese Orthotrichia species are provided.

ZooKeys, Feb 10, 2012

Four new species of genus Plectrocnemia and 4 new species of genus Nyctiophylax are described, na... more Four new species of genus Plectrocnemia and 4 new species of genus Nyctiophylax are described, namely: Plectrocnemia verticalis sp. n.; Plectrocnemia acuminata sp. n.; Plectrocnemia cryptoparamere sp. n.; Plectrocnemia qianshanensis sp. n.; Nyctiophylax (Nyctiophylax) senticosus sp. n.; Nyctiophylax (Paranyctiophylax) gracilis sp. n.; Nyctiophylax (Paranyctiophylax) pungens sp. n.; and Nyctiophylax (Paranyctiophylax) auriculatus sp. n.

Zoosymposia, Jun 10, 2011

The 13th International Symposium on Trichoptera took place in Bialowieża, Poland, in 22-27 June 2... more The 13th International Symposium on Trichoptera took place in Bialowieża, Poland, in 22-27 June 2009, but the idea to organize the symposium there was evoked 3 years earlier. When, at the end of the symposium in Mexico City, Professor Hans Malicky suggested that the next symposium take place in Poland, the first question we asked ourselves was, "Where could we organize it?" That night we were discussing this issue with several colleagues and Wolfram Mey said, "Why not do it in Białowieża?" The idea was tempting. In the last 2 decades Trichoptera symposia were organized in big cities. The last one even took place in one of the largest cities of the world-Mexico City. By contrast, Białowieża is just a small village; but for researchers it is paradise. The significance of this last natural European forest for science is reflected in the presence of many scientific institutions there. And so, we decided that Bialowieża would be a perfect venue for our symposium. Three years latter in June 2009, 65 Trichoptera researchers from 24 countries met here. Over 35 oral presentations and 41 posters were delivered. All of them were of high quality and many of them are published in this volume. An important part of the symposium was various trips and excursions organized for the participants and accompanying persons. Although summer is usually warm and sunny in Poland, this year proved to be exceptionally wet and stormy. One such powerful thunderstorm descended on Bialowieża Forest in the midst of a walking tour of the Protected Area of the National Park. Within minutes all of us were soaked. We walked quickly through the primeval forest in the near darkness, interrupted occasionally by frightening flashes of lightening. While not entirely pleasant, this hike made us realize the power of the natural elements. The accompanying persons enjoyed trips in the surrounding area, visiting picturesque villages hidden among gently rolling hills. The caddis on the symposium logo is Tricholeiochiton fagesii (Guinard), a species with a small larva that has extremely long legs and a transparent, secreted case. It is rare in Poland and very beautiful. We were inspired by a picture from Bert Higler 's book, "De Nederlandse Kokerjufferlarven," in which a group of larvae looks similar to galloping horses. Bert actively participated in most of our symposia and wanted very much to be with us in Bialowieża. Unfortunately, due to illness he was unable to attend. A few months later Bert passed away. We dedicate this volume to his memory. Proceedings of previous international symposia on Trichoptera have been published by a variety of publishers, including Dr.

Journal of entomology and zoology studies, Jun 1, 2015

Information on the aquatic insects and their benefits and risks to the society are scanty among t... more Information on the aquatic insects and their benefits and risks to the society are scanty among the general public, students and the scientific community, when compared with the same on the land forms. In this article, an attempt is made to overcome this deficiency. A brief description is furnished along with the representative photographs of eleven orders of aquatic insects. These orders are Collembola (springtails), Ephemeroptera (mayflies), Odonata (dragonflies and damselflies), Plecoptera (stoneflies), Hemiptera (true bugs), Megaloptera (dobsonflies and alderflies), Neuroptera (songillaflies), Trichoptera (caddisflies), Lepidoptera (butterflies and moths), Coleoptera (beetles) and Diptera (true flies). Detailed information is presented on the beneficial role of aquatic insects in food webs, biomonitoring, fishing and control of noxious weeds. The harmful impacts caused by these animals to the society and the ecosystem by way of general nuisance, transmission of diseases and destruction of crops, are described. The importance of the need for a new generation of aquatic entomologists, is stressed.

Journal of Entomological Science, Oct 1, 1997

Journal of The North American Benthological Society, Dec 1, 1994

Annals of the Entomological Society of America, 1981

Zoosymposia, Jun 12, 2020

Genetic sequencing (with mtCoI) was used to associate larvae with two new Vietnamese caddisfly (t... more Genetic sequencing (with mtCoI) was used to associate larvae with two new Vietnamese caddisfly (trichoptera) species from Bach Mã National Park, Hydromanicus calyx n. sp. (hydropsychidae) and Drepanocentron dentatum n. sp. (Xiphocentronidae). Adult Drepanocentron dentatum n. sp. is distinguishable by the toothy dorsal margin of the male inferior appendages; the most similar species, D. vercaius, has teeth on the ventral, but not the dorsal margins of the inferior appendages. the larva of D. dentatum is fully described herein, the first larva to be described in the genus Drepanocentron; it can be separated from the other two known xiphocentronid larvae (Xiphocentron messapus and Cnodocentron yavapai) by the smooth, thin mesal margin of each mandible. Adult Hydromanicus calyx n. sp. is distinguishable by the cupped apices of the male harpagones; this character is absent in morphologically similar species (H. nieuwenhuisi, H. abiud, and H. serubabel). the larva of H. calyx is morphologically most similar to that of H. inferior, but can be distinguished by the presence of faint muscle scars interrupting the lateral margins of the stridulatory areas on the venter of the head.

Beiträge zur Entomologie, Dec 7, 2023

Zootaxa, 2019

Four new species of genus Rhyacophila are described and illustrated from Thailand and Myanmar, So... more Four new species of genus Rhyacophila are described and illustrated from Thailand and Myanmar, Southeast Asia. Rhyacophila longicaudata sp. n. is in the R. nigrocephala Group; the very long basal segment of each inferior appendage distinguishes it from other related species. Rhyacophila aksornkoaei sp. n. and R. limsakuli sp. n. are in the R. anatina Group: they can be differentiated from other species of the group by the brush-like parameres and presence of a ventral process of the aedeagus in R. aksornkoaei sp. n., and by the rectangular apical segment of each inferior appendage and the hooked parameres in R. limsakuli sp. n.. Rhyacophila kengtungensis is in the R. yishepa Group and is characterized by the subtriangular shape of the preanal appendages in dorsal view and by the very large dorsal appendages of the phallic apparatus.

Beiträge zur Entomologie, Dec 21, 2023

Based on a recalibrated BEAST diversification time analysis, we provide a revised chronology for ... more Based on a recalibrated BEAST diversification time analysis, we provide a revised chronology for the evolution of major lineages of Trichoptera. Fossil evidence indicates that caddisflies evolved at least by the Norian of Late Triassic (median age 222.6 Ma), compared with our estimate of at least 201.3 Ma. The ancestors of suborders Annulipalpia and Integripalpia also evolved as early as the Norian. Fossil evidence indicates that the ancestor of subterorder Phryganides lived at least by the Aalenian of Middle Jurassic (median age 173.6 Ma), compared with our estimate of at least 174.1 Ma.

Insecta Mundi, Sep 1, 1997

The phylogenetic relationships of the species ofPadunielia are analyzed based on characters oflar... more The phylogenetic relationships of the species ofPadunielia are analyzed based on characters oflarvae, pupae, and adults (mainly male genitalia). The genus is monophyletic and most closely related to Psychomyia, and Metalype in the subfamily Psychornyiinae. Nine species groups are suggested. Eight species, including six new to science, are reported from the People's Republic of China for the first time.

Zoosymposia, Jun 12, 2020

Taxonomy and Basic Description Protoptila morettii was described by John Morse in 1990 from male ... more Taxonomy and Basic Description Protoptila morettii was described by John Morse in 1990 from male and female adults collected in South Carolina. Males measure 3.2 to 3.5 mm (0.13 to 0.14 inches) from front of head to tips of folded wings, with forewings each 2.7 to 3.0 mm (0.11 to 0.12 inches) long. Females measure 3.1 to 4.1mm (0.12 to 0.16 inches) from front of head to tips of folded wings, with forewings each 2.9 to 3.8 mm (0.11 to 0.15 inches) long. The eggs, larvae and pupae of this species remain unknown. Species of Protoptila are members of the subfamily Protoptilinae, family Glossosomatidae, order Trichoptera. The Trichoptera, or caddisflies, are holometabolous insects that are most closely related to Lepidoptera, or moths. Trichoptera eggs, larvae and pupae usually are fully submerged in water and depend on oxygenated water for respiration. Adult caddisflies are aerial/terrestrial, but are rarely found far from water. Adults generally resemble those of their Lepidoptera cousins except, rather than being covered by scales, Trichoptera wings are typically clothed with hair, which is presumably important in repelling water, hence the Latin name trichos (hairy) and ptera (wings). Trichoptera larvae generally resemble those of Lepidoptera except that caddisflies never have fleshy, crochet-bearing prolegs on middle abdominal segments.

Zoosymposia, Jun 12, 2020

Zootaxa, Sep 14, 2017

The Chinese species of the genera Lannapsyche and Marilia (Trichoptera, Odontoceridae) are review... more The Chinese species of the genera Lannapsyche and Marilia (Trichoptera, Odontoceridae) are reviewed, including 1 new species of Lannapsyche (Lannapsyche altilis sp. n.), 1 new Chinese record of Lannapsyche (Lannapsyche chantaramongkolae Malicky 1989, originally described from Thailand), and 3 new species of Marilia (Marilia discaulis, sp. n., Marilia megalopos, sp. n., and Marilia qinlingensis, sp. n.), bringing the Chinese fauna of Lannapsyche to 3 species and of Marilia to 6 species. Previously known species Lannapsyche setschuana Malicky 1995b and Marilia parallela Hwang 1957 are redescribed and reillustrated for both males and females. Updated information about the holotype specimen of Marilia simulans Forsslund 1935 is provided. Distinctive differences of the female genitalia are described, which may add important diagnostic characters for genus and species recognition.

Zenodo (CERN European Organization for Nuclear Research), Dec 31, 2012

Currently, 21 species of the genus Plectrocnemia are known from China. Examination of material co... more Currently, 21 species of the genus Plectrocnemia are known from China. Examination of material collected from 13 provinces of China from 1990-2005 has revealed 9 new species and 3 new records of this genus, bringing the number of Chinese Plectrocnemia species to 33. Newly described species include:

Zoosymposia, Jun 12, 2020

Zootaxa, Aug 1, 2014

Currently, 8 species of the genus Nyctiophylax Brauer are known from China. Examination of materi... more Currently, 8 species of the genus Nyctiophylax Brauer are known from China. Examination of material collected from Guangdong, Guangxi, Jiangxi and Sichuan Provinces during 2004-2005 has revealed 4 new species and 2 new records of this genus, bringing the number of Chinese Nyctiophylax species to 14. Newly described species include: Nyctiophylax (Paranyctiophylax) crinalis n. sp., N. (P.) dactylatus n. sp., N. (P.) orbicularis n. sp., and N. (P.) macrorrhinus n. sp. Nyctiophylax (Paranyctiophylax) sagax Mey and N. (N.) amphonion Malicky & Chantaramongkol are newly recorded for the Chinese fauna, which are re-illustrated and re-described for clear comparisons. The additional collection sites for the previously described species are provided.

Zenodo (CERN European Organization for Nuclear Research), Dec 31, 2010

Six new species of the caddisfly genus Orthotrichia are described and illustrated from China, nam... more Six new species of the caddisfly genus Orthotrichia are described and illustrated from China, namely Or. apophysis Zhou & Yang, sp. nov.; Or. latiramifera Zhou & Yang, sp. nov.; Or. cornuta Zhou &Yang, sp. nov.; Or. discedata Zhou & Morse, sp. nov.; Or. cuspidigera Zhou & Yang, sp. nov.; and Or. subrhomba Zhou & Morse, sp. nov.. One species, Or. terpsichore Malicky & Chantaramongkol, 2007, is recorded for the first time from China, bringing the number of species of the genus known from China to 14. Of these, 10 species are distributed among 3 species groups of Marshall (1979), with 4 species unassigned. The known distributions for all the Chinese Orthotrichia species are provided.

ZooKeys, Feb 10, 2012

Four new species of genus Plectrocnemia and 4 new species of genus Nyctiophylax are described, na... more Four new species of genus Plectrocnemia and 4 new species of genus Nyctiophylax are described, namely: Plectrocnemia verticalis sp. n.; Plectrocnemia acuminata sp. n.; Plectrocnemia cryptoparamere sp. n.; Plectrocnemia qianshanensis sp. n.; Nyctiophylax (Nyctiophylax) senticosus sp. n.; Nyctiophylax (Paranyctiophylax) gracilis sp. n.; Nyctiophylax (Paranyctiophylax) pungens sp. n.; and Nyctiophylax (Paranyctiophylax) auriculatus sp. n.

Zoosymposia, Jun 10, 2011

The 13th International Symposium on Trichoptera took place in Bialowieża, Poland, in 22-27 June 2... more The 13th International Symposium on Trichoptera took place in Bialowieża, Poland, in 22-27 June 2009, but the idea to organize the symposium there was evoked 3 years earlier. When, at the end of the symposium in Mexico City, Professor Hans Malicky suggested that the next symposium take place in Poland, the first question we asked ourselves was, "Where could we organize it?" That night we were discussing this issue with several colleagues and Wolfram Mey said, "Why not do it in Białowieża?" The idea was tempting. In the last 2 decades Trichoptera symposia were organized in big cities. The last one even took place in one of the largest cities of the world-Mexico City. By contrast, Białowieża is just a small village; but for researchers it is paradise. The significance of this last natural European forest for science is reflected in the presence of many scientific institutions there. And so, we decided that Bialowieża would be a perfect venue for our symposium. Three years latter in June 2009, 65 Trichoptera researchers from 24 countries met here. Over 35 oral presentations and 41 posters were delivered. All of them were of high quality and many of them are published in this volume. An important part of the symposium was various trips and excursions organized for the participants and accompanying persons. Although summer is usually warm and sunny in Poland, this year proved to be exceptionally wet and stormy. One such powerful thunderstorm descended on Bialowieża Forest in the midst of a walking tour of the Protected Area of the National Park. Within minutes all of us were soaked. We walked quickly through the primeval forest in the near darkness, interrupted occasionally by frightening flashes of lightening. While not entirely pleasant, this hike made us realize the power of the natural elements. The accompanying persons enjoyed trips in the surrounding area, visiting picturesque villages hidden among gently rolling hills. The caddis on the symposium logo is Tricholeiochiton fagesii (Guinard), a species with a small larva that has extremely long legs and a transparent, secreted case. It is rare in Poland and very beautiful. We were inspired by a picture from Bert Higler 's book, "De Nederlandse Kokerjufferlarven," in which a group of larvae looks similar to galloping horses. Bert actively participated in most of our symposia and wanted very much to be with us in Bialowieża. Unfortunately, due to illness he was unable to attend. A few months later Bert passed away. We dedicate this volume to his memory. Proceedings of previous international symposia on Trichoptera have been published by a variety of publishers, including Dr.

Journal of entomology and zoology studies, Jun 1, 2015

Information on the aquatic insects and their benefits and risks to the society are scanty among t... more Information on the aquatic insects and their benefits and risks to the society are scanty among the general public, students and the scientific community, when compared with the same on the land forms. In this article, an attempt is made to overcome this deficiency. A brief description is furnished along with the representative photographs of eleven orders of aquatic insects. These orders are Collembola (springtails), Ephemeroptera (mayflies), Odonata (dragonflies and damselflies), Plecoptera (stoneflies), Hemiptera (true bugs), Megaloptera (dobsonflies and alderflies), Neuroptera (songillaflies), Trichoptera (caddisflies), Lepidoptera (butterflies and moths), Coleoptera (beetles) and Diptera (true flies). Detailed information is presented on the beneficial role of aquatic insects in food webs, biomonitoring, fishing and control of noxious weeds. The harmful impacts caused by these animals to the society and the ecosystem by way of general nuisance, transmission of diseases and destruction of crops, are described. The importance of the need for a new generation of aquatic entomologists, is stressed.

Journal of Entomological Science, Oct 1, 1997

Journal of The North American Benthological Society, Dec 1, 1994

Annals of the Entomological Society of America, 1981

Zoosymposia, Jun 12, 2020

Genetic sequencing (with mtCoI) was used to associate larvae with two new Vietnamese caddisfly (t... more Genetic sequencing (with mtCoI) was used to associate larvae with two new Vietnamese caddisfly (trichoptera) species from Bach Mã National Park, Hydromanicus calyx n. sp. (hydropsychidae) and Drepanocentron dentatum n. sp. (Xiphocentronidae). Adult Drepanocentron dentatum n. sp. is distinguishable by the toothy dorsal margin of the male inferior appendages; the most similar species, D. vercaius, has teeth on the ventral, but not the dorsal margins of the inferior appendages. the larva of D. dentatum is fully described herein, the first larva to be described in the genus Drepanocentron; it can be separated from the other two known xiphocentronid larvae (Xiphocentron messapus and Cnodocentron yavapai) by the smooth, thin mesal margin of each mandible. Adult Hydromanicus calyx n. sp. is distinguishable by the cupped apices of the male harpagones; this character is absent in morphologically similar species (H. nieuwenhuisi, H. abiud, and H. serubabel). the larva of H. calyx is morphologically most similar to that of H. inferior, but can be distinguished by the presence of faint muscle scars interrupting the lateral margins of the stridulatory areas on the venter of the head.

Beiträge zur Entomologie, Dec 7, 2023

Zootaxa, 2019

Four new species of genus Rhyacophila are described and illustrated from Thailand and Myanmar, So... more Four new species of genus Rhyacophila are described and illustrated from Thailand and Myanmar, Southeast Asia. Rhyacophila longicaudata sp. n. is in the R. nigrocephala Group; the very long basal segment of each inferior appendage distinguishes it from other related species. Rhyacophila aksornkoaei sp. n. and R. limsakuli sp. n. are in the R. anatina Group: they can be differentiated from other species of the group by the brush-like parameres and presence of a ventral process of the aedeagus in R. aksornkoaei sp. n., and by the rectangular apical segment of each inferior appendage and the hooked parameres in R. limsakuli sp. n.. Rhyacophila kengtungensis is in the R. yishepa Group and is characterized by the subtriangular shape of the preanal appendages in dorsal view and by the very large dorsal appendages of the phallic apparatus.