Özgür KOÇAK | Cukurova University (original) (raw)

Papers by Özgür KOÇAK

Temnostethus (Temnostethus) gracilis Hor... more Temnostethus (Temnostethus) gracilis Horváth, 1907 (Fig. 5G) Material examined. TURKEY: İSTANBUL: Esenyurt, 41°03′01.7″N 28°40′34.2″E, 16.v.2018, 1 3, B. Çerçi leg.and det. (BCIT). Collection circumstances. It was collected while it was wandering on a rock under a Pyrus sp. (Rosaceae). Comment. Temnostethus gracilis is a widespread species known from most of Europe, including the Balkans. It is a predacious species known to feed on Sternorrhyncha species. It was observed to live on numerous deciduous tree species such as Malus, Pyrus, Prunus, Crataegus (Rosaceae), Fraxinus (Oleaceae), Quercus, Fagus (Fagaceae), Betula, Corylus (Betulaceae), Acer (Sapindaceae), Salix, Populus (Salicaceae) (PÉRICART 1972). Temnostethus gracilis differs from T. winkelmanni WAGNER, 1961 and T. longirostris (Horváth, 1907) in shorter clypeus and the existence of brachypterous form, from T. pusillus (Herrich-Schäffer, 1835) by pale second antennal segment, and from T. tibialis Reuter, 1888 by much shorter and sparser setae on hemelytra which are almost invisible (Fig. 5G) (PÉRICART 1972). Distribution in Turkey. İstanbul (this work).

Hallodapus concolor (Reuter, 1890) (Fig. 5D) Material examined. TURKE... more Hallodapus concolor (Reuter, 1890) (Fig. 5D) Material examined. TURKEY: İZMIR: Urla, 38°18′33.9″N 26°43′47.8″E: 17.vii.2018, 2 33; 22.vii.2018, 1 3; 2.viii.2018, 1 3, B. Çerçi leg.and det.(BCIT). Collecting circumstances. Light trap. Comment. The genus Hallodapus Fieber, 1858 is represented by two species in Turkey: H. suturalis (Herrich-Schäffer, 1839) and H. pseudoconcolor (Linnavuori, 1984). While the former is widely distributed in the West Palaearctic Region, the latter is only known from Iraq and the South East of Turkey (MATOCQ et al. 2014). Hallodapus concolor Reuter, 1890 is distributed in tropical Africa from Cameroon to Sudan and in Asia from Saudi Arabia and Azerbaijan all the way to Kirghizia and in Europe only in Crete (Greece) (KERZHNER & JOSIFOV 1999). This species was observed to live on low grass vegetation in dry sandy areas (LINNAVUORI 1996). Hallodapus concolor (Figs 8A, B) is most similar to H. pseudoconcolor but differs from it in the shape of the left paramere and theca (see LINNAVUORI 1984). Distribution in Turkey. İzmir (this work).

Fig. 5. A ‒ Phytocoris (Exophytocoris) carapezzai sp. nov., alive female attracted to UV light; B... more Fig. 5. A ‒ Phytocoris (Exophytocoris) carapezzai sp. nov., alive female attracted to UV light; B‒C ‒ Campylomma miyamotoi Yasunaga, 2001 (B ‒ male; C ‒ female). D ‒ Hallodapus concolor (Reuter, 1890), male; E ‒ Zanchius breviceps (Wagner, 1951), alive male on Ficus sp.; F ‒ Montandoniola moraguesi (Puton, 1896), alive male attracted to UV light; G ‒ Temnostethus (Temnostethus) gracilis Horváth, 1907, male; H ‒ Plinthisus (Isioscytus) minutissimus Fieber, 1864, female.

Fig. 4. A‒B ‒ Maurodactylus kukuensis V. G. Putshkov, 1978: A ‒ male; B ‒ female. C ‒ Pinalitus v... more Fig. 4. A‒B ‒ Maurodactylus kukuensis V. G. Putshkov, 1978: A ‒ male; B ‒ female. C ‒ Pinalitus viscicola (Puton, 1888), male. D ‒ Platycranus (Genistocapsus) alkestis Linnavuori, 1999, male. Scale bars: 1 mm.

Fig. 3. A‒B ‒ Campylomma miyamotoi Yasunaga, 2001: A ‒ male, B ‒ vesica. C ‒ Compsidolon (Compsid... more Fig. 3. A‒B ‒ Campylomma miyamotoi Yasunaga, 2001: A ‒ male, B ‒ vesica. C ‒ Compsidolon (Compsidolon) elegantulum Reuter, 1899, male. Scale bars: A, C ‒ 1 mm; B ‒ 0.1 mm.

FIG. 8—Orthotylus (Pinocapsus) kmenti Çerçi & Koçak sp. n., holotype, male, dorsal habitus (Scale... more FIG. 8—Orthotylus (Pinocapsus) kmenti Çerçi & Koçak sp. n., holotype, male, dorsal habitus (Scale bar = 1 mm).

FIG. 10–33. 10–17—Vesical processes, 18–33—Left and right parameres. 10, 18, 19—O. (P.) fuscescen... more FIG. 10–33. 10–17—Vesical processes, 18–33—Left and right parameres. 10, 18, 19—O. (P.) fuscescens (Kirschbaum, 1856), 11, 20, 21—O. (P.) cupressi Reuter, 1883, 12, 22, 23—O. (P.) callitris Lindberg, 1940, 13, 24, 25—O. (P.) thaleia Linnavuori, 1999, 14, 26, 27—O. (P.) alashanensis Tian & Nonnaizab, 1999, 15, 28, 29—O. (P.) sabinae Tian & Nonnaizab, 1999, 16, 30, 31—O. (P.) gemmae Gessé & Goula, 2003, 17, 32, 33—O. (P.) olympicus Matocq, Pagola-Carte & Pluot-Sigwalt, 2018 [Fig. 10, 18, 19 adopted from (Wyniger & Burckhardt 2003); Fig. 11, 12, 20, 21, 22, 23 adopted from (Wagner 1974); Fig. 13, 24, 25 adopted from (Linnavuori 1999); Fig. 14, 15, 26, 27, 28, 29 adopted from (Tian & Nonnaizab 1999); Fig. 16, 30, 31 adopted from (Gessé & Goula 2003); Fig. 17, 32, 33 adopted from (Matocq et al. 2018)]. (Scale bars = 0.1 mm if not stated otherwise).

FIG. 4A–C. 4A—Fulvius oxycarenoides (Reuter, 1878), dorsal habitus (scale bar = 1 mm), 4B—idem, h... more FIG. 4A–C. 4A—Fulvius oxycarenoides (Reuter, 1878), dorsal habitus (scale bar = 1 mm), 4B—idem, head and pronotum from lateral view (Scale bar = 0.1 mm), 4C—idem, pronotum from superolateral view.

Zenodo (CERN European Organization for Nuclear Research), May 31, 2023

A list of 414 species belonging to 30 families of Heteroptera from Karaman, Türkiye is compiled, ... more A list of 414 species belonging to 30 families of Heteroptera from Karaman, Türkiye is compiled, based on previous records in literature and new records from comprehensive field studies conducted in numerous localities between 2010 and 2021. Among them 254 species are recorded from Karaman for the first time, Adelphophylus pericarti Matocq & Magnien, 2009, Teratocoris antennatus (Boheman, 1852) and Dicyphus (Dicyphus) lindbergi Wagner, 1951 (Miridae) constitute new records for the fauna of Türkiye. Additionally, 41 species are recorded for the first time, either from Central Anatolian or Mediterranean regions. A chorotype analysis is done to illustrate the main elements of the Heteroptera fauna of Karaman province. This chorotype analysis revealed that Heteroptera fauna of Karaman is mainly composed of Mediterranean related species (32%, 128 spp.), followed by Middle Eastern related (16%, 66 spp.) and European related (15%, 62 spp.) species. The number of Anatolian endemic species is 25, four of which are endemic to Karaman. These findings indicate that the Heteroptera fauna of Karaman has remained mostly undiscovered until now and although present study is a major step forward in understanding the species diversity of this region, future studies are needed to uncover the true biodiversity of Karaman province.

TURKISH JOURNAL OF ZOOLOGY, 2021

The Harlequin Ladybird, Harmonia axyridis (Pallas, 1773), is native to Eastern Asia but it has be... more The Harlequin Ladybird, Harmonia axyridis (Pallas, 1773), is native to Eastern Asia but it has been introduced to many countries in Africa, America, and Eurasia. In Turkey, H. axyridis is one of the most fast-spreading invasive alien species. Since the collection of the first specimens in 2010, H. axyridis spread almost all over the country. The species spreads fast in Turkey according to the new records, obtained originally from the field studies and the citizen science data. It was recorded for the first time from 19 of 81 Turkish provinces

A new species of Dorcadion Dalman, 1817 from Turkey (Coleoptera: Cerambycidae), 2012

The following new taxon is described: Dorcadion (Cribridorcadion) karamanense sp. n. from Karaman... more The following new taxon is described: Dorcadion (Cribridorcadion) karamanense sp. n. from Karaman province (Turkey), close to D. oezdurali Önalp, 1988.

Fig. 2. Phytocoris (Exophytocoris) carapezzai sp. nov.: A ‒ male; B ‒ posterior leg; C‒D ‒ vesica... more Fig. 2. Phytocoris (Exophytocoris) carapezzai sp. nov.: A ‒ male; B ‒ posterior leg; C‒D ‒ vesica in two different views; E ‒ left paramere; F ‒ right paramere. Scale bars: A, B ‒ 1 mm; C, D ‒ 0.1 mm; E, F ‒ 0.1 mm.

Phytocoris (Exophytocoris) carapezzai sp... more Phytocoris (Exophytocoris) carapezzai sp. nov. (Figs 2, 5A) Type locality. Turkey, Karaman, Merkez, 37°13′18.3″N 33°13′14.2″E. Material examined. HOLOTYPE: 1 3 (AZMM), TURKEY: KARAMAN: Merkez, 37°13′18.3″N 33°13′14.2″E, 8.ix.2015; glued on a pointed cardboard with male genitalia on the same cardbord, labelled as follows: ʻTurkey, Karaman prov. / Merkez, with UV light trap / Ö. Koçak coll. [white printed label] // Holotypus / Phytocoris (Exophytocoris) carapezzai sp. n. / B. Çerçi det. 2019 [red printed label]'. PARATYPES: TURKEY: KARAMAN: 2 33, Merkez, 8.ix.2015, light trap, Ö. Koçak leg., B. Çerçi det. (BCIT). Diagnosis. The new species is recognized by the combination of the following characters: general coloration yellowish brown, pronotum darker than hemelytra and scutellum. First antennal segment white with dense red patterns, eyes very big, ocular index 0.6–0.8 in male. Posterior margin of pronotum with narrow white marginal band and wavy and uninterrupted brown submarginal band. Hemelytra with scattered reddish pattern, tip of clavus darkened, outer margin of corium and inner margin of cuneus with red dots interrupted by white coloration along their whole length, inner upper corner of cuneus with isolated red dot. Membrane brownish and translucent with dense small pale patches. Femora (Fig. 2B) dark brown with dense white dots. Left paramere edentate (Fig. 2E), hypophysis strongly enlarged before apex, vesica with marginally dentate lamellae apically (Fig. 2D), sclerotized process of vesica (Fig. 2C) stick-like, long and thick, edentate and slightly bent towards apex. Description. Male. Coloration. General coloration yellowish brown (Fig. 2A). Head yellowish brown with reddish patterns. First antennal segment whitish, maculated in red, second antennal segment yellow except for white basal ring, third antennal segment black with white ring both basally and apically, apical ring sometimes absent, last segment unicoloro [...]

Two subspecies of Dorcadion Dalman, 1817 from Turkey (Coleoptera: Cerambycidae), 2016

The following new subspecies are described: Dorcadion menradi pilosicollis ssp. nov. from Karaman... more The following new subspecies are described: Dorcadion menradi pilosicollis ssp. nov. from Karaman province, and Dorcadion lameerei konyaense ssp. nov. from Konya province.

In this study six Heteroptera species are reported for the first time from Turkey; Dicyphus (Meso... more In this study six Heteroptera species are reported for the first time from Turkey; Dicyphus (Mesodiyphus) martinoi Josifov, 1958; Campylomma simillimum Jakovlev, 1882; Orthotylus (Melanotrichus) rubidus (Puton 1874); Orthotylus (Litocoris) ericetorum arboreae Wagner, 1969; Heegeria tangirica (Saunders, 1877) and Brachysteles parvicornis (Costa, 1847). Additionally a new synonymy is proposed; Orius (Heterorius) laticollis laticollis (Reuter, 1884) = Orius (Heterorius) laticollis discolor (Reuter, 1884) syn. n. and a remarkable color form of C. simillimum is mentioned.

A total of 11 new Heteroptera species are recorded from Turkey for the first time: Amphiareus con... more A total of 11 new Heteroptera species are recorded from Turkey for the first time: Amphiareus constrictus (Stal, 1860), Anthocoris simulans (Reuter, 1884), Halyomorpha halys ( Stal , 1855), Engistus exsanguis (Stal, 1872), Campylomma unicolor Poppius, 1914, Campylomma vendicarinum Carapezza, 1991, Capsodes gothicus (Linnaeus, 1758), Leucodellus zagdani (Putshkov, 1970), Phytocoris scitulus scitulus Reuter, 1908, Psallus cruentatus (Mulsant and Rey, 1852) and Solenoxyphus punctipennis (Reuter, 1879). Additionally, Phytocoris (Phytocoris) malckyi Rieger, 1995 is synonymized with Phytocoris (Exophytocoris) parvuloides Wagner, 1961; Phytocoris (Exophytocoris) parvuloides = Phytocoris (Phytocoris) malckyi Rieger, 1995 syn. nov.

Montandoniola moraguesi (Puton, 1896) (Fig. 5F) Material examined: TU... more Montandoniola moraguesi (Puton, 1896) (Fig. 5F) Material examined: TURKEY: MERSIN: Anamur, 36°02′08.3″N 32°48′31.7″E, 17.vii.2018, 13, Ö. Koçak leg., B. Çerçi det.The specimen was only photographed but not collected. Comment. Montandoniola moraguesi is an important biological control agent used to fight thrips invasions on economically important crops. It is unique among the Anthocoridae species known from Turkey because of its thrips-like appearance (Fig.11B). It lives inside the galls or gall-like deformations caused by the thrips on which it feeds (PLUOT- SIGWALT et al. 2009). Montandoniola moraguesi is known from the following countries: the Balearic Islands, France, Italy, Spain, the Canary Islands, Israel, Morocco, Algeria, Tunisia, Egypt, Chad, Sudan, Burkina Faso, and South Africa (PLUOT- SIGWALT et al. 2009). Considering its wide distribution along the Mediterranean coasts, it is possible that this species has a stable population in Mersin, Turkey and was not introduced here by farmers. Distribution in Turkey. Mersin (this work).

Two subspecies of Dorcadion infernale Mulsant & Rey, 1863 from Turkey (Coleoptera: Cerambycidae), 2016

The following new subspecies are described: Dorcadion infernale costatidorsum ssp. nov. from Anka... more The following new subspecies are described: Dorcadion infernale costatidorsum ssp. nov. from Ankara province, and Dorcadion infernale luteosutura ssp. nov. from Karaman province.

Journal of Insect Biodiversity, 2016

This study is based on material of the suborder Heteroptera collected and photographed from diffe... more This study is based on material of the suborder Heteroptera collected and photographed from different provinces of Turkey between 2011 and 2015. In this study, 15 species belonging to 9 families are recorded from various provinces of Turkey. All of the species listed include remarks on their known distribution either in Turkey or in worldwide. Among them 12 species are new for the Heteroptera fauna of Turkey. In addition it is found out that Pasira marinadolina Putshkov & Moulet, 2003 could also have bicolored connexivum.

Zenodo (CERN European Organization for Nuclear Research), May 31, 2023

TURKISH JOURNAL OF ZOOLOGY, 2021

A new species of Dorcadion Dalman, 1817 from Turkey (Coleoptera: Cerambycidae), 2012

Two subspecies of Dorcadion Dalman, 1817 from Turkey (Coleoptera: Cerambycidae), 2016

Two subspecies of Dorcadion infernale Mulsant & Rey, 1863 from Turkey (Coleoptera: Cerambycidae), 2016

Journal of Insect Biodiversity, 2016

Karaman ili, Kazımkarabekir ilçesi sınırları içerisinde yer alan Hacıbaba Dağı, Torosların, İç An... more Karaman ili, Kazımkarabekir ilçesi sınırları içerisinde yer alan Hacıbaba Dağı, Torosların, İç Anadolu bölgesindeki en kuzey yükseltisidir. Dağın zirvesi Hacıbaba tepesi olarak anılır ve 2450 m. dir. Dağ, karbonatlı kayaçlardan oluşmuştur. Doğu kısmında yoğun metamorfizma ve tektonizma etkileri görülür. Bazı bölgelerinde siyah mermer için ocaklar açılmış ve halen işletilmektedir. Genel olarak bitki örtüsünü meşe ormanları oluşturur. 1500 m. üzerinde ise ardıç ağaçları boy göstermeye başlar. Dağın birçok bölgesi üzüm bağlarıyla kaplıdır. Şu anda hastalık ve bakımsızlıktan büyük bölümü terkedilmiş bu bağlar kelebekler için çok ideal yaşam alanlar oluştururlar. Dağı oluşturan kayaçların genel eğimi güney doğrultulu olduğu için kuzey cephesinde su kaynakları çok azdır. Buna rağmen kelebek açısından kuzey bakı çok daha zengindir. Gerek tür çeşitliliği gerek popülasyonlar çok yoğundur. Bölgede ilkbahar aylarında rengarenk olan doğa haziran ortalarına doğru sarı rengin hakimiyetine bürünür. Ama yinede adaçayları, kekikler ve sığırkuyrukları sonbahar aylarının ortalarına kadar çiçeklidir. Ağustos aylarında yol kenarlarında yoğunlaşan deve dikenleri, su kıyılarındaki yabani nane ve yavşanlar özellikle kelebekler için adeta bir çekim merkezi haline gelirler. Bölgede kelebek gözlemi This watermark does not appear in the registered version