Frank-Thorsten Krell | Denver Museum of Nature and Science (original) (raw)
Papers by Frank-Thorsten Krell
The Bulletin of zoological nomenclature, Jul 30, 2020
Introduction: In the Afrotropics, insect assemblages on carrion have barely been studied. After l... more Introduction: In the Afrotropics, insect assemblages on carrion have barely been studied. After large scavengers have removed the majority of meat and internal organs, the remaining carcass is mainly consumed by scarab beetles and flies. Methods: From 1995 to 1998 we studied the beetle component of necrocenoses in the Parc National de la Comoé, in northeastern Ivory Coast. Mainly using fresh goat skins, we set up about 70 baits which were exposed for one to seven weeks in savanna parkland, gallery forest, and enclosed grassland of the river valley; some baits were shaded, some watered, and there was an untreated control. We collected all beetles from the baits three times a day resulting in over 60,000 specimens over the course of this study. Results/Conclusion: Scarabaeoidea make up 99.7% of the necrophilous beetle fauna with Phaeochrous (Hybosoridae) being the most abundant group, followed by Trogidae. Differences of beetle activities between times of day and between habitats, respectively, could clearly be demonstrated. The main factor determining abundance of necrophilous beetles are single, large rainfall events. On the days after heavy rainfall the baits were visited by numbers of beetles often magnitudes larger than during dry periods. Watering the baits or offering high-quality (fresh) bait could not replicate this effect.
The Bulletin of zoological nomenclature, Apr 30, 2021
BioOne Complete (complete.BioOne.org) is a full-text database of 200 subscribed and open-access t... more BioOne Complete (complete.BioOne.org) is a full-text database of 200 subscribed and open-access titles in the biological, ecological, and environmental sciences published by nonprofit societies, associations, museums, institutions, and presses.
Entomologische Nachrichten und Berichte, Sep 19, 1996
Bulletin of Zoological Nomenclature, 2008
Chemoecology, Mar 3, 2017
compounds) to investigate which components or simple mixtures are cues for several taxa of dung b... more compounds) to investigate which components or simple mixtures are cues for several taxa of dung beetles. Additionally, we analyzed the degree of specialization of dung beetle species and communities on particular scent types and herbivore dung. Our results show that butyric acid in particular is an important volatile cue for dung beetles. Dung beetles show a preference for some scent types, but turned out to be generalists. This finding is in congruence with the assumption that organisms living from ephemeral resources should rather be generalists instead of specialists.
Zenodo (CERN European Organization for Nuclear Research), Dec 31, 2016
The mango-flower beetle, Protaetia fusca (Coleoptera: Scarabaeidae: Cetoniinae) is newly recorded... more The mango-flower beetle, Protaetia fusca (Coleoptera: Scarabaeidae: Cetoniinae) is newly recorded for Wake Island. The distribution and invasive history of this species on tropical oceanic islands are briefly reviewed and the causes for its success as an established alien and sometimes invasive species discussed.
Systematic Entomology, 2004
Deutsche Entomologische Zeitschrift, Sep 4, 1992
Alpine Entomology, Jul 26, 2018
On distribution and morphology of some Onthophagus species in Switzerland (Coleoptera: Scarabaeid... more On distribution and morphology of some Onthophagus species in Switzerland (Coleoptera: Scarabaeidae).-On the basis of 1097 specimens from 12 collections, the distribution of ten dung beetle species of the genus Onthophagus (Scarabaeidae) in Switzerland
The Bulletin of zoological nomenclature, 2007
The Bulletin of zoological nomenclature, Aug 1, 2017
The purpose of this application, under Articles 70.2 and 80 of the Code, is to conserve the curre... more The purpose of this application, under Articles 70.2 and 80 of the Code, is to conserve the current and universal usage of the genus name Heteronychus Dejean, 1833 for a genus of rhinoceros beetles (Scarabaeidae: Dynastinae). Several species in this genus are important pests in Africa, Asia and Australia. The correct, long-overlooked type species is Geotrupes syrichtus Fabricius, 1775, designated by Duponchel (1845), is currently recognized as a valid species and the type species of Syrictes Prell, 1935, a genus placed in a different tribe of Dynastinae. Acceptance of this type species would transfer a widely used name from an economically important to a rarely cited genus, and the economically important genus would need to be named by the scarcely used subjective junior synonym Heteronychidius Paulian, 1954 with a dubious taxonomic identity. To conserve the usage of the widely used name Heteronychus, it is proposed that all type fixations for this genus preceding that of Heteronychus cricetus Hausmann, 1807 by Krell (2002) be set aside.
The Bulletin of zoological nomenclature, Oct 15, 2022
BioOne Complete (complete.BioOne.org) is a full-text database of 200 subscribed and open-access t... more BioOne Complete (complete.BioOne.org) is a full-text database of 200 subscribed and open-access titles in the biological, ecological, and environmental sciences published by nonprofit societies, associations, museums, institutions, and presses.
Zootaxa, Nov 25, 2015
Hemicyrthus blaffarti new species (Coleoptera: Scarabaeidae: Dynastinae) from the Parc Provincial... more Hemicyrthus blaffarti new species (Coleoptera: Scarabaeidae: Dynastinae) from the Parc Provincial de la Rivière Bleue in southern New Caledonia is described. It differs from the most similar species, H. elongatus, by the completely and sharply margined pronotal base, the presence of elytral micropunctures, shorter apical setae of the mesotibiae and metatibiae, and a different shape of the parameres. For the first time, a sexually dimorphic character is described in this genus: the median emargination of the last sternite has a developed margin in males, but not in females. A revised key for all Hemicyrthus species is given. Hemicyrthus as a genus of short-range endemics, and its biogeographical relationships are briefly discussed.
Zootaxa, Nov 12, 2021
The scientific literature shows no consistency in attributing the authorship of the name for the ... more The scientific literature shows no consistency in attributing the authorship of the name for the dor beetle, Anoplotrupes stercorosus (Coleoptera: Geotrupidae: Geotrupinae: Geotrupini). The species is widespread all over Europe, Siberia, Turkey (Nikolajev et al. 2016), and Iran (Samin et al. 2018), feeding on dung, carrion, mushrooms, mouldy litter, heaps of marc, and tree sap (Koch [1991]: 351; Byk & Semkiw 2010). Most authors who dealt with the taxon credited its authorship to Ludwig Gottlieb Scriba, the author of the paper in which the taxon was described, while few others stated that Hartmann-cited by Scriba himself in the same paper-is responsible for the name. There has been no agreement even on the publication year of the original description, although the majority of authors dated it to 1791. The aim of this paper is to establish once and for all the authorship of the specific name and the publication date of the paper in which the taxon was first described. Scarabaeus stercorosus was first described in Scriba's article "Entomologische Bemerkungen und Erfahrungen", included in the third issue of the first volume of Scriba's own magazine "Journal für die Liebhaber der Entomologie" (Fig. 1). However, Scriba integrated text from Hartmann in the form of long quotes (see below) into that paper. As far as we know, the first author who assigned authorship of the name to Hartmann alone in a scientific publication was Hoppe (1795) who wrote on page 9, "Scarabaeus stercorosus Hartm.". Previously, the reporter who wrote a review of the third issue of Scriba's journal for the "Allgemeine Literatur-Zeitung", an exhaustive magazine on German and European cultural production at the time, had spoken of the "Hartmannsche Scarabaeus stercorosus" [Hartmann's Scarabaeus stercorosus] (Anonymous 1792). During the 19 th century the specific name stercorosus, as Geotrupes stercorosus, was credited to Hartmann by only a few authors, such as Mulsant (1842), Erichson (1848), Murray (1853), Mulsant & Rey (1871), and Bertolini (1872), whereas the vast majority of authors assigned the authorship of the name to Scriba. This situation had not changed until the end of the 20 th century. The authorship of stercorosus, first still in Geotrupes, later as Anoplotrupes stercorosus, continued to be assigned to Scriba, at least until the 1980s, when some authors, par
The Bulletin of zoological nomenclature, 2002
Zoology in The Middle East, Jul 20, 2021
Occurrence records of the genus Orubesa Reitter, 1895, in the Arabian Penisula and surrounding ar... more Occurrence records of the genus Orubesa Reitter, 1895, in the Arabian Penisula and surrounding areas are presented. Morphological study of types and additional specimens, for the first time considering endophallic sclerites, revealed that all northeast African and Arabian specimens belong to one taxon, Orubesa plicifrons semenowi (Arrow, 1911). The synonymy of both O. sudanica (Balthasar, 1871) and O. linnavuorii (Endrödi, 1970) with O. plicifrons semenowi is confirmed. The first records of Orubesa from Yemen are given. http://www.zoobank.org/urn:lsid:zoobank.org:pub:62C7A1C5-1669-42A3-B840-FFC6E098221E
The Bulletin of zoological nomenclature, 2007
The purpose of this application, under Article 23.9.3 of the Code, is to conserve the specific na... more The purpose of this application, under Article 23.9.3 of the Code, is to conserve the specific name of the dung beetle Onthophagus sycophanta Fairmaire, 1887 from China and Vietnam. This name is a junior primary homonym of the forgotten name Onthophagus sycophanta Mulsant, 1842 from Europe (Coleoptera, SCARABAEIDAE). O. sycophanta Fairmaire, 1887 has been in regular use since its description whereas O. sycophanta Mulsant, 1842 has never been used after its description. It is proposed to conserve the name O. sycophanta Fairmaire, 1887 by suppression of the name O. sycophanta Mulsant, 1842.
Coleopterists Bulletin, Jun 27, 2017
The Bulletin of zoological nomenclature, Jul 30, 2020
Introduction: In the Afrotropics, insect assemblages on carrion have barely been studied. After l... more Introduction: In the Afrotropics, insect assemblages on carrion have barely been studied. After large scavengers have removed the majority of meat and internal organs, the remaining carcass is mainly consumed by scarab beetles and flies. Methods: From 1995 to 1998 we studied the beetle component of necrocenoses in the Parc National de la Comoé, in northeastern Ivory Coast. Mainly using fresh goat skins, we set up about 70 baits which were exposed for one to seven weeks in savanna parkland, gallery forest, and enclosed grassland of the river valley; some baits were shaded, some watered, and there was an untreated control. We collected all beetles from the baits three times a day resulting in over 60,000 specimens over the course of this study. Results/Conclusion: Scarabaeoidea make up 99.7% of the necrophilous beetle fauna with Phaeochrous (Hybosoridae) being the most abundant group, followed by Trogidae. Differences of beetle activities between times of day and between habitats, respectively, could clearly be demonstrated. The main factor determining abundance of necrophilous beetles are single, large rainfall events. On the days after heavy rainfall the baits were visited by numbers of beetles often magnitudes larger than during dry periods. Watering the baits or offering high-quality (fresh) bait could not replicate this effect.
The Bulletin of zoological nomenclature, Apr 30, 2021
BioOne Complete (complete.BioOne.org) is a full-text database of 200 subscribed and open-access t... more BioOne Complete (complete.BioOne.org) is a full-text database of 200 subscribed and open-access titles in the biological, ecological, and environmental sciences published by nonprofit societies, associations, museums, institutions, and presses.
Entomologische Nachrichten und Berichte, Sep 19, 1996
Bulletin of Zoological Nomenclature, 2008
Chemoecology, Mar 3, 2017
compounds) to investigate which components or simple mixtures are cues for several taxa of dung b... more compounds) to investigate which components or simple mixtures are cues for several taxa of dung beetles. Additionally, we analyzed the degree of specialization of dung beetle species and communities on particular scent types and herbivore dung. Our results show that butyric acid in particular is an important volatile cue for dung beetles. Dung beetles show a preference for some scent types, but turned out to be generalists. This finding is in congruence with the assumption that organisms living from ephemeral resources should rather be generalists instead of specialists.
Zenodo (CERN European Organization for Nuclear Research), Dec 31, 2016
The mango-flower beetle, Protaetia fusca (Coleoptera: Scarabaeidae: Cetoniinae) is newly recorded... more The mango-flower beetle, Protaetia fusca (Coleoptera: Scarabaeidae: Cetoniinae) is newly recorded for Wake Island. The distribution and invasive history of this species on tropical oceanic islands are briefly reviewed and the causes for its success as an established alien and sometimes invasive species discussed.
Systematic Entomology, 2004
Deutsche Entomologische Zeitschrift, Sep 4, 1992
Alpine Entomology, Jul 26, 2018
On distribution and morphology of some Onthophagus species in Switzerland (Coleoptera: Scarabaeid... more On distribution and morphology of some Onthophagus species in Switzerland (Coleoptera: Scarabaeidae).-On the basis of 1097 specimens from 12 collections, the distribution of ten dung beetle species of the genus Onthophagus (Scarabaeidae) in Switzerland
The Bulletin of zoological nomenclature, 2007
The Bulletin of zoological nomenclature, Aug 1, 2017
The purpose of this application, under Articles 70.2 and 80 of the Code, is to conserve the curre... more The purpose of this application, under Articles 70.2 and 80 of the Code, is to conserve the current and universal usage of the genus name Heteronychus Dejean, 1833 for a genus of rhinoceros beetles (Scarabaeidae: Dynastinae). Several species in this genus are important pests in Africa, Asia and Australia. The correct, long-overlooked type species is Geotrupes syrichtus Fabricius, 1775, designated by Duponchel (1845), is currently recognized as a valid species and the type species of Syrictes Prell, 1935, a genus placed in a different tribe of Dynastinae. Acceptance of this type species would transfer a widely used name from an economically important to a rarely cited genus, and the economically important genus would need to be named by the scarcely used subjective junior synonym Heteronychidius Paulian, 1954 with a dubious taxonomic identity. To conserve the usage of the widely used name Heteronychus, it is proposed that all type fixations for this genus preceding that of Heteronychus cricetus Hausmann, 1807 by Krell (2002) be set aside.
The Bulletin of zoological nomenclature, Oct 15, 2022
BioOne Complete (complete.BioOne.org) is a full-text database of 200 subscribed and open-access t... more BioOne Complete (complete.BioOne.org) is a full-text database of 200 subscribed and open-access titles in the biological, ecological, and environmental sciences published by nonprofit societies, associations, museums, institutions, and presses.
Zootaxa, Nov 25, 2015
Hemicyrthus blaffarti new species (Coleoptera: Scarabaeidae: Dynastinae) from the Parc Provincial... more Hemicyrthus blaffarti new species (Coleoptera: Scarabaeidae: Dynastinae) from the Parc Provincial de la Rivière Bleue in southern New Caledonia is described. It differs from the most similar species, H. elongatus, by the completely and sharply margined pronotal base, the presence of elytral micropunctures, shorter apical setae of the mesotibiae and metatibiae, and a different shape of the parameres. For the first time, a sexually dimorphic character is described in this genus: the median emargination of the last sternite has a developed margin in males, but not in females. A revised key for all Hemicyrthus species is given. Hemicyrthus as a genus of short-range endemics, and its biogeographical relationships are briefly discussed.
Zootaxa, Nov 12, 2021
The scientific literature shows no consistency in attributing the authorship of the name for the ... more The scientific literature shows no consistency in attributing the authorship of the name for the dor beetle, Anoplotrupes stercorosus (Coleoptera: Geotrupidae: Geotrupinae: Geotrupini). The species is widespread all over Europe, Siberia, Turkey (Nikolajev et al. 2016), and Iran (Samin et al. 2018), feeding on dung, carrion, mushrooms, mouldy litter, heaps of marc, and tree sap (Koch [1991]: 351; Byk & Semkiw 2010). Most authors who dealt with the taxon credited its authorship to Ludwig Gottlieb Scriba, the author of the paper in which the taxon was described, while few others stated that Hartmann-cited by Scriba himself in the same paper-is responsible for the name. There has been no agreement even on the publication year of the original description, although the majority of authors dated it to 1791. The aim of this paper is to establish once and for all the authorship of the specific name and the publication date of the paper in which the taxon was first described. Scarabaeus stercorosus was first described in Scriba's article "Entomologische Bemerkungen und Erfahrungen", included in the third issue of the first volume of Scriba's own magazine "Journal für die Liebhaber der Entomologie" (Fig. 1). However, Scriba integrated text from Hartmann in the form of long quotes (see below) into that paper. As far as we know, the first author who assigned authorship of the name to Hartmann alone in a scientific publication was Hoppe (1795) who wrote on page 9, "Scarabaeus stercorosus Hartm.". Previously, the reporter who wrote a review of the third issue of Scriba's journal for the "Allgemeine Literatur-Zeitung", an exhaustive magazine on German and European cultural production at the time, had spoken of the "Hartmannsche Scarabaeus stercorosus" [Hartmann's Scarabaeus stercorosus] (Anonymous 1792). During the 19 th century the specific name stercorosus, as Geotrupes stercorosus, was credited to Hartmann by only a few authors, such as Mulsant (1842), Erichson (1848), Murray (1853), Mulsant & Rey (1871), and Bertolini (1872), whereas the vast majority of authors assigned the authorship of the name to Scriba. This situation had not changed until the end of the 20 th century. The authorship of stercorosus, first still in Geotrupes, later as Anoplotrupes stercorosus, continued to be assigned to Scriba, at least until the 1980s, when some authors, par
The Bulletin of zoological nomenclature, 2002
Zoology in The Middle East, Jul 20, 2021
Occurrence records of the genus Orubesa Reitter, 1895, in the Arabian Penisula and surrounding ar... more Occurrence records of the genus Orubesa Reitter, 1895, in the Arabian Penisula and surrounding areas are presented. Morphological study of types and additional specimens, for the first time considering endophallic sclerites, revealed that all northeast African and Arabian specimens belong to one taxon, Orubesa plicifrons semenowi (Arrow, 1911). The synonymy of both O. sudanica (Balthasar, 1871) and O. linnavuorii (Endrödi, 1970) with O. plicifrons semenowi is confirmed. The first records of Orubesa from Yemen are given. http://www.zoobank.org/urn:lsid:zoobank.org:pub:62C7A1C5-1669-42A3-B840-FFC6E098221E
The Bulletin of zoological nomenclature, 2007
The purpose of this application, under Article 23.9.3 of the Code, is to conserve the specific na... more The purpose of this application, under Article 23.9.3 of the Code, is to conserve the specific name of the dung beetle Onthophagus sycophanta Fairmaire, 1887 from China and Vietnam. This name is a junior primary homonym of the forgotten name Onthophagus sycophanta Mulsant, 1842 from Europe (Coleoptera, SCARABAEIDAE). O. sycophanta Fairmaire, 1887 has been in regular use since its description whereas O. sycophanta Mulsant, 1842 has never been used after its description. It is proposed to conserve the name O. sycophanta Fairmaire, 1887 by suppression of the name O. sycophanta Mulsant, 1842.
Coleopterists Bulletin, Jun 27, 2017