Curtis Marean - Academia.edu (original) (raw)

Papers by Curtis Marean

Research paper thumbnail of Size variation in Tachyoryctes splendens (East African mole-rat) and its implications for late Quaternary temperature change in equatorial East Africa

This study develops a new proxy for Quaternary temperature change in tropical Africa through anal... more This study develops a new proxy for Quaternary temperature change in tropical Africa through analysis of size variation in East African mole-rat (Tachyoryctes splendens). In modern mole-rats, mandibular alveolar length is unrelated to annual precipitation, precipitation seasonality, temperature seasonality, or primary productivity. However, it is inversely correlated with mean annual temperature, in agreement with Bergmann's rule. This relationship is observed at temperatures below ∼17.3 °C, but not at higher temperatures. We apply these observations to late Quaternary mole-rats from Wakondo (∼100 ka) and Kisaaka (∼50 ka) in the Lake Victoria region and Enkapune ya Muto (EYM; ∼7.2–3.2 ka) in Kenya's central rift. The Lake Victoria mole-rats are larger than expected for populations from warm climates typical of the area today, implying cooler temperatures in the past. The magnitude of temperature decline needed to drive the size shift is substantial (∼4–6 °C), similar in magn...

Research paper thumbnail of Paleolithic hominin remains from Eshkaft-e Gavi (southern Zagros Mountains, Iran): description, affinities, and evidence for butchery

Eshkaft-e Gavi is a cave located in the southern Zagros Mountains of Iran and is one of the few a... more Eshkaft-e Gavi is a cave located in the southern Zagros Mountains of Iran and is one of the few archaeological sites in the region to preserve both Middle Paleolithic and Upper Paleolithic occupations. Excavation of the site in the 1970s yielded an assemblage of lithic and faunal remains, including ten hominin specimens: a mandibular molar, four cranial fragments, a clavicular diaphysis, the proximal half of a metacarpal, a fragment of os coxa, the proximal diaphysis of a juvenile femur, and a patella. The bones derive from a minimum of four individuals, including two juveniles. Although many of these remains could be Epi-Paleolithic in age, one of the juvenile specimens-the mandibular molar-occurs at the base of the cave's Upper Paleolithic sequence. The remains are very fragmentary, but those that preserve diagnostic morphology indicate that they represent modern humans. The molar is taxonomically diagnostic, thus confirming the association of the Aurignacian-like Baradostian ...

Research paper thumbnail of The stratigraphy of the Middle Stone Age sediments at Pinnacle Point Cave 13B (Mossel Bay, Western Cape Province, South Africa)

Pinnacle Point Cave 13B (PP13B) has provided the earliest archaeological evidence for the exploit... more Pinnacle Point Cave 13B (PP13B) has provided the earliest archaeological evidence for the exploitation of marine shellfish, along with very early evidence for use and modification of pigments and the production of bladelets, all dated to approximately 164 ka (Marean et al., 2007). This makes PP13B a key site in studies of the origins of modern humans, one of a handful of sites in Africa dating to Marine Isotope Stage 6 (MIS 6), and the only site on the coast of South Africa with human occupation confidently dated to MIS 6. Along with this MIS 6 occupation there are rich archaeological sediments dated to MIS 5, and together these sediments are differentially preserved in three different areas of the cave. The sediments represent a complex palimpsest of geogenic, biogenic, and anthropogenic input and alteration that are described and interpreted through the use of a variety of macrostratigraphic, micromorphologic, and geochemical techniques. Three independent dating techniques allow u...

Research paper thumbnail of A worked bone assemblage from 120,000–90,000 year old deposits at Contrebandiers Cave, Atlantic Coast, Morocco

Research paper thumbnail of Cave life histories of non-anthropogenic sediments help us understand associated archaeological contexts

Quaternary Research

Pinnacle Point (PP) near Mossel Bay in the Western Cape Province, South Africa, is known for a se... more Pinnacle Point (PP) near Mossel Bay in the Western Cape Province, South Africa, is known for a series of archaeological caves with important archaeological finds. Extensive excavations and studies in two of them (PP13B and PP5-6) have documented alternating periods of anthropogenic-dominated and geogenic-dominated sedimentation. A variety of caves do not bear evidence of anthropogenic remains. We have studied in detail the remnant deposits of three of them, Staircase Cave, Crevice Cave, and PP29, which have been formed under the same geologic and sedimentary conditions with those with anthropogenic contributions. Their remains are small and patchy but have extensive speleothem formations (as do most caves at PP) that were isotopically analyzed for paleoclimate and paleoenvironmental reconstruction. These caves also offer the opportunity to understand the purely geogenic signature of the PP locality and thus offer a geogenic baseline for the anthropogenic caves. Archaeologists normal...

Research paper thumbnail of The Palaeo-Agulhas Plain: Temporal and spatial variation in an extraordinary extinct ecosystem of the Pleistocene of the Cape Floristic Region

Quaternary Science Reviews

Research paper thumbnail of Seasonal scheduling of shellfish collection in the Middle and Later Stone Ages of southern Africa

Journal of Human Evolution

Research paper thumbnail of Seasonal scheduling of shellfish collection in the Middle and Later Stone Ages of southern Africa

Journal of Human Evolution, 2019

This study assesses the seasonal scheduling of shellfish harvesting among hunter-gatherer populat... more This study assesses the seasonal scheduling of shellfish harvesting among hunter-gatherer populations along the southernmost coast of South Africa, based on a large number of serial oxygen isotope analyses of marine mollusk shells from four archaeological sites. The south coast of South Africa boasts an exceptional record of coastal hunter-gatherer occupation spanning the Holocene, the last glacial cycle and beyond. The significance of coastal adaptations, in this region in particular, for later modern human evolution has been prominently debated. Shellfishing behaviors are an important focus for investigation given the dietary and scheduling implications and the abundant archaeological shell remains in numerous sites. Key to better understanding coastal foraging is whether it was limited to one particular season, or year-round. Yet, this has proven very difficult to establish by conventional archaeological methods. This study reconstructs seasonal harvesting patterns by calculating...

Research paper thumbnail of Palaeoecology of giraffe tracks in Late Pleistocene aeolianites on the Cape south coast

South African Journal of Science

Until now there have been no reliable historical or skeletal fossil records for the giraffe (Gira... more Until now there have been no reliable historical or skeletal fossil records for the giraffe (Giraffa camelopardalis) south of the Orange River or northern Namaqualand. The recent discovery of fossil giraffe tracks in coastal aeolianites east of Still Bay, South Africa, significantly increases the geographical range for this species, and has implications for Late Pleistocene climate and vegetation in the southern Cape. Giraffe populations have specialised needs, and require a savanna ecosystem. Marine geophysical and geological evidence suggests that the broad, currently submerged floodplains of the Gouritz and Breede Rivers likely supported a productive savanna of Vachellia karroo during Pleistocene glacial conditions, which would have provided a suitable habitat for this species. We show evidence for the hypothesis that the opening of the submerged shelf during glacial periods acted as a pathway for mammals to migrate along the southern coastal plain.

Research paper thumbnail of Phytoliths as an indicator of early modern humans plant gathering strategies, fire fuel and site occupation intensity during the Middle Stone Age at Pinnacle Point 5-6 (south coast, South Africa)

PloS one, 2018

The study of plant remains in archaeological sites, along with a better understanding of the use ... more The study of plant remains in archaeological sites, along with a better understanding of the use of plants by prehistoric populations, can help us shed light on changes in survival strategies of hunter-gatherers and consequent impacts on modern human cognition, social organization, and technology. The archaeological locality of Pinnacle Point (Mossel Bay, South Africa) includes a series of coastal caves, rock-shelters, and open-air sites with human occupations spanning the Acheulian through Middle Stone Age (MSA) and Later Stone Age (LSA). These sites have provided some of the earliest evidence for complex human behaviour and technology during the MSA. We used phytoliths-amorphous silica particles that are deposited in cells of plants-as a proxy for the reconstruction of past human plant foraging strategies on the south coast of South Africa during the Middle and Late Pleistocene, emphasizing the use and control of fire as well as other possible plant uses. We analysed sediment samp...

Research paper thumbnail of The trajectory of bone surface modification studies in paleoanthropology and a new Bayesian solution to the identification controversy

Journal of human evolution, 2017

A critical issue in human evolution is how to determine when hominins began incorporating signifi... more A critical issue in human evolution is how to determine when hominins began incorporating significant amounts of meat into their diets. This fueled evolution of a larger brain and other adaptations widely considered unique to modern humans. Determination of the spatiotemporal context of this shift rests on accurate identification of fossil bone surface modifications (BSM), such as stone tool butchery marks. Multidecade-long debates over the agents responsible for individual BSM are indicative of systemic flaws in current approaches to identification. Here we review the current state of BSM studies and introduce a novel probabilistic approach to identifying agents of BSM. We use control assemblages of bones modified by modern agents to train a multivariate Bayesian probability model. The model then identifies BSM agents with associated uncertainties, serving as the basis for a predictive probabilistic algorithm. The multivariate Bayesian approach offers a novel, probabilistic, and an...

Research paper thumbnail of Further human fossils from the Middle Stone Age deposits of Die Kelders Cave 1, Western Cape Province, South Africa

Journal of human evolution, 2017

Research paper thumbnail of Chronometric investigations of the Middle to Upper Paleolithic transition in the Zagros Mountains using AMS radiocarbon dating and Bayesian age modelling

Journal of human evolution, 2017

The Middle to Upper Paleolithic transition is often linked with a bio-cultural shift involving th... more The Middle to Upper Paleolithic transition is often linked with a bio-cultural shift involving the dispersal of modern humans outside of Africa, the concomitant replacement of Neanderthals across Eurasia, and the emergence of new technological traditions. The Zagros Mountains region assumes importance in discussions concerning this period as its geographic location is central to all pertinent hominin migration areas, pointing to both east and west. As such, establishing a reliable chronology in the Zagros Mountains is crucial to our understanding of these biological and cultural developments. Political circumstance, coupled with the poor preservation of organic material, has meant that a clear chronological definition of the Middle to Upper Paleolithic transition for the Zagros Mountains region has not yet been achieved. To improve this situation, we have obtained new archaeological samples for AMS radiocarbon dating from three sites: Kobeh Cave, Kaldar Cave, and Ghār-e Boof (Iran)....

Research paper thumbnail of Lithic technological responses to Late Pleistocene glacial cycling at Pinnacle Point Site 5-6, South Africa

PloS one, 2017

There are multiple hypotheses for human responses to glacial cycling in the Late Pleistocene, inc... more There are multiple hypotheses for human responses to glacial cycling in the Late Pleistocene, including changes in population size, interconnectedness, and mobility. Lithic technological analysis informs us of human responses to environmental change because lithic assemblage characteristics are a reflection of raw material transport, reduction, and discard behaviors that depend on hunter-gatherer social and economic decisions. Pinnacle Point Site 5-6 (PP5-6), Western Cape, South Africa is an ideal locality for examining the influence of glacial cycling on early modern human behaviors because it preserves a long sequence spanning marine isotope stages (MIS) 5, 4, and 3 and is associated with robust records of paleoenvironmental change. The analysis presented here addresses the question, what, if any, lithic assemblage traits at PP5-6 represent changing behavioral responses to the MIS 5-4-3 interglacial-glacial cycle? It statistically evaluates changes in 93 traits with no a priori as...

Research paper thumbnail of Foraging potential of underground storage organ plants in the southern Cape, South Africa

Journal of Human Evolution, 2016

Research paper thumbnail of Protein sequences bound to mineral surfaces persist into deep time

eLife, Jan 27, 2016

Proteins persist longer in the fossil record than DNA, but the longevity, survival mechanisms and... more Proteins persist longer in the fossil record than DNA, but the longevity, survival mechanisms and substrates remain contested. Here, we demonstrate the role of mineral binding in preserving the protein sequence in ostrich (Struthionidae) eggshell, including from the palaeontological sites of Laetoli (3.8 Ma) and Olduvai Gorge (1.3 Ma) in Tanzania. By tracking protein diagenesis back in time we find consistent patterns of preservation, demonstrating authenticity of the surviving sequences. Molecular dynamics simulations of struthiocalcin-1 and -2, the dominant proteins within the eggshell, reveal that distinct domains bind to the mineral surface. It is the domain with the strongest calculated binding energy to the calcite surface that is selectively preserved. Thermal age calculations demonstrate that the Laetoli and Olduvai peptides are 50 times older than any previously authenticated sequence (equivalent to ~16 Ma at a constant 10°C).

Research paper thumbnail of R. Dale Guthrie: Frozen Fauna of the Mammoth Steppe: The Story of Blue Babe

J Vertebrate Paleontol, 1990

Research paper thumbnail of Hunter-Gatherer Foraging Strategies in Tropical Grasslands: Model Building and Testing in the East African Middle and Later Stone Age

Journal of Anthropological Archaeology, Sep 1, 1997

Research paper thumbnail of Large mammal skeletal element transport: applying foraging theory in a complex taphonomic system

Journal of Taphonomy, 2003

Research paper thumbnail of Later quaternary extinct ungulates of East Africa and palaeoenvironmental implications

Research paper thumbnail of Size variation in Tachyoryctes splendens (East African mole-rat) and its implications for late Quaternary temperature change in equatorial East Africa

This study develops a new proxy for Quaternary temperature change in tropical Africa through anal... more This study develops a new proxy for Quaternary temperature change in tropical Africa through analysis of size variation in East African mole-rat (Tachyoryctes splendens). In modern mole-rats, mandibular alveolar length is unrelated to annual precipitation, precipitation seasonality, temperature seasonality, or primary productivity. However, it is inversely correlated with mean annual temperature, in agreement with Bergmann's rule. This relationship is observed at temperatures below ∼17.3 °C, but not at higher temperatures. We apply these observations to late Quaternary mole-rats from Wakondo (∼100 ka) and Kisaaka (∼50 ka) in the Lake Victoria region and Enkapune ya Muto (EYM; ∼7.2–3.2 ka) in Kenya's central rift. The Lake Victoria mole-rats are larger than expected for populations from warm climates typical of the area today, implying cooler temperatures in the past. The magnitude of temperature decline needed to drive the size shift is substantial (∼4–6 °C), similar in magn...

Research paper thumbnail of Paleolithic hominin remains from Eshkaft-e Gavi (southern Zagros Mountains, Iran): description, affinities, and evidence for butchery

Eshkaft-e Gavi is a cave located in the southern Zagros Mountains of Iran and is one of the few a... more Eshkaft-e Gavi is a cave located in the southern Zagros Mountains of Iran and is one of the few archaeological sites in the region to preserve both Middle Paleolithic and Upper Paleolithic occupations. Excavation of the site in the 1970s yielded an assemblage of lithic and faunal remains, including ten hominin specimens: a mandibular molar, four cranial fragments, a clavicular diaphysis, the proximal half of a metacarpal, a fragment of os coxa, the proximal diaphysis of a juvenile femur, and a patella. The bones derive from a minimum of four individuals, including two juveniles. Although many of these remains could be Epi-Paleolithic in age, one of the juvenile specimens-the mandibular molar-occurs at the base of the cave's Upper Paleolithic sequence. The remains are very fragmentary, but those that preserve diagnostic morphology indicate that they represent modern humans. The molar is taxonomically diagnostic, thus confirming the association of the Aurignacian-like Baradostian ...

Research paper thumbnail of The stratigraphy of the Middle Stone Age sediments at Pinnacle Point Cave 13B (Mossel Bay, Western Cape Province, South Africa)

Pinnacle Point Cave 13B (PP13B) has provided the earliest archaeological evidence for the exploit... more Pinnacle Point Cave 13B (PP13B) has provided the earliest archaeological evidence for the exploitation of marine shellfish, along with very early evidence for use and modification of pigments and the production of bladelets, all dated to approximately 164 ka (Marean et al., 2007). This makes PP13B a key site in studies of the origins of modern humans, one of a handful of sites in Africa dating to Marine Isotope Stage 6 (MIS 6), and the only site on the coast of South Africa with human occupation confidently dated to MIS 6. Along with this MIS 6 occupation there are rich archaeological sediments dated to MIS 5, and together these sediments are differentially preserved in three different areas of the cave. The sediments represent a complex palimpsest of geogenic, biogenic, and anthropogenic input and alteration that are described and interpreted through the use of a variety of macrostratigraphic, micromorphologic, and geochemical techniques. Three independent dating techniques allow u...

Research paper thumbnail of A worked bone assemblage from 120,000–90,000 year old deposits at Contrebandiers Cave, Atlantic Coast, Morocco

Research paper thumbnail of Cave life histories of non-anthropogenic sediments help us understand associated archaeological contexts

Quaternary Research

Pinnacle Point (PP) near Mossel Bay in the Western Cape Province, South Africa, is known for a se... more Pinnacle Point (PP) near Mossel Bay in the Western Cape Province, South Africa, is known for a series of archaeological caves with important archaeological finds. Extensive excavations and studies in two of them (PP13B and PP5-6) have documented alternating periods of anthropogenic-dominated and geogenic-dominated sedimentation. A variety of caves do not bear evidence of anthropogenic remains. We have studied in detail the remnant deposits of three of them, Staircase Cave, Crevice Cave, and PP29, which have been formed under the same geologic and sedimentary conditions with those with anthropogenic contributions. Their remains are small and patchy but have extensive speleothem formations (as do most caves at PP) that were isotopically analyzed for paleoclimate and paleoenvironmental reconstruction. These caves also offer the opportunity to understand the purely geogenic signature of the PP locality and thus offer a geogenic baseline for the anthropogenic caves. Archaeologists normal...

Research paper thumbnail of The Palaeo-Agulhas Plain: Temporal and spatial variation in an extraordinary extinct ecosystem of the Pleistocene of the Cape Floristic Region

Quaternary Science Reviews

Research paper thumbnail of Seasonal scheduling of shellfish collection in the Middle and Later Stone Ages of southern Africa

Journal of Human Evolution

Research paper thumbnail of Seasonal scheduling of shellfish collection in the Middle and Later Stone Ages of southern Africa

Journal of Human Evolution, 2019

This study assesses the seasonal scheduling of shellfish harvesting among hunter-gatherer populat... more This study assesses the seasonal scheduling of shellfish harvesting among hunter-gatherer populations along the southernmost coast of South Africa, based on a large number of serial oxygen isotope analyses of marine mollusk shells from four archaeological sites. The south coast of South Africa boasts an exceptional record of coastal hunter-gatherer occupation spanning the Holocene, the last glacial cycle and beyond. The significance of coastal adaptations, in this region in particular, for later modern human evolution has been prominently debated. Shellfishing behaviors are an important focus for investigation given the dietary and scheduling implications and the abundant archaeological shell remains in numerous sites. Key to better understanding coastal foraging is whether it was limited to one particular season, or year-round. Yet, this has proven very difficult to establish by conventional archaeological methods. This study reconstructs seasonal harvesting patterns by calculating...

Research paper thumbnail of Palaeoecology of giraffe tracks in Late Pleistocene aeolianites on the Cape south coast

South African Journal of Science

Until now there have been no reliable historical or skeletal fossil records for the giraffe (Gira... more Until now there have been no reliable historical or skeletal fossil records for the giraffe (Giraffa camelopardalis) south of the Orange River or northern Namaqualand. The recent discovery of fossil giraffe tracks in coastal aeolianites east of Still Bay, South Africa, significantly increases the geographical range for this species, and has implications for Late Pleistocene climate and vegetation in the southern Cape. Giraffe populations have specialised needs, and require a savanna ecosystem. Marine geophysical and geological evidence suggests that the broad, currently submerged floodplains of the Gouritz and Breede Rivers likely supported a productive savanna of Vachellia karroo during Pleistocene glacial conditions, which would have provided a suitable habitat for this species. We show evidence for the hypothesis that the opening of the submerged shelf during glacial periods acted as a pathway for mammals to migrate along the southern coastal plain.

Research paper thumbnail of Phytoliths as an indicator of early modern humans plant gathering strategies, fire fuel and site occupation intensity during the Middle Stone Age at Pinnacle Point 5-6 (south coast, South Africa)

PloS one, 2018

The study of plant remains in archaeological sites, along with a better understanding of the use ... more The study of plant remains in archaeological sites, along with a better understanding of the use of plants by prehistoric populations, can help us shed light on changes in survival strategies of hunter-gatherers and consequent impacts on modern human cognition, social organization, and technology. The archaeological locality of Pinnacle Point (Mossel Bay, South Africa) includes a series of coastal caves, rock-shelters, and open-air sites with human occupations spanning the Acheulian through Middle Stone Age (MSA) and Later Stone Age (LSA). These sites have provided some of the earliest evidence for complex human behaviour and technology during the MSA. We used phytoliths-amorphous silica particles that are deposited in cells of plants-as a proxy for the reconstruction of past human plant foraging strategies on the south coast of South Africa during the Middle and Late Pleistocene, emphasizing the use and control of fire as well as other possible plant uses. We analysed sediment samp...

Research paper thumbnail of The trajectory of bone surface modification studies in paleoanthropology and a new Bayesian solution to the identification controversy

Journal of human evolution, 2017

A critical issue in human evolution is how to determine when hominins began incorporating signifi... more A critical issue in human evolution is how to determine when hominins began incorporating significant amounts of meat into their diets. This fueled evolution of a larger brain and other adaptations widely considered unique to modern humans. Determination of the spatiotemporal context of this shift rests on accurate identification of fossil bone surface modifications (BSM), such as stone tool butchery marks. Multidecade-long debates over the agents responsible for individual BSM are indicative of systemic flaws in current approaches to identification. Here we review the current state of BSM studies and introduce a novel probabilistic approach to identifying agents of BSM. We use control assemblages of bones modified by modern agents to train a multivariate Bayesian probability model. The model then identifies BSM agents with associated uncertainties, serving as the basis for a predictive probabilistic algorithm. The multivariate Bayesian approach offers a novel, probabilistic, and an...

Research paper thumbnail of Further human fossils from the Middle Stone Age deposits of Die Kelders Cave 1, Western Cape Province, South Africa

Journal of human evolution, 2017

Research paper thumbnail of Chronometric investigations of the Middle to Upper Paleolithic transition in the Zagros Mountains using AMS radiocarbon dating and Bayesian age modelling

Journal of human evolution, 2017

The Middle to Upper Paleolithic transition is often linked with a bio-cultural shift involving th... more The Middle to Upper Paleolithic transition is often linked with a bio-cultural shift involving the dispersal of modern humans outside of Africa, the concomitant replacement of Neanderthals across Eurasia, and the emergence of new technological traditions. The Zagros Mountains region assumes importance in discussions concerning this period as its geographic location is central to all pertinent hominin migration areas, pointing to both east and west. As such, establishing a reliable chronology in the Zagros Mountains is crucial to our understanding of these biological and cultural developments. Political circumstance, coupled with the poor preservation of organic material, has meant that a clear chronological definition of the Middle to Upper Paleolithic transition for the Zagros Mountains region has not yet been achieved. To improve this situation, we have obtained new archaeological samples for AMS radiocarbon dating from three sites: Kobeh Cave, Kaldar Cave, and Ghār-e Boof (Iran)....

Research paper thumbnail of Lithic technological responses to Late Pleistocene glacial cycling at Pinnacle Point Site 5-6, South Africa

PloS one, 2017

There are multiple hypotheses for human responses to glacial cycling in the Late Pleistocene, inc... more There are multiple hypotheses for human responses to glacial cycling in the Late Pleistocene, including changes in population size, interconnectedness, and mobility. Lithic technological analysis informs us of human responses to environmental change because lithic assemblage characteristics are a reflection of raw material transport, reduction, and discard behaviors that depend on hunter-gatherer social and economic decisions. Pinnacle Point Site 5-6 (PP5-6), Western Cape, South Africa is an ideal locality for examining the influence of glacial cycling on early modern human behaviors because it preserves a long sequence spanning marine isotope stages (MIS) 5, 4, and 3 and is associated with robust records of paleoenvironmental change. The analysis presented here addresses the question, what, if any, lithic assemblage traits at PP5-6 represent changing behavioral responses to the MIS 5-4-3 interglacial-glacial cycle? It statistically evaluates changes in 93 traits with no a priori as...

Research paper thumbnail of Foraging potential of underground storage organ plants in the southern Cape, South Africa

Journal of Human Evolution, 2016

Research paper thumbnail of Protein sequences bound to mineral surfaces persist into deep time

eLife, Jan 27, 2016

Proteins persist longer in the fossil record than DNA, but the longevity, survival mechanisms and... more Proteins persist longer in the fossil record than DNA, but the longevity, survival mechanisms and substrates remain contested. Here, we demonstrate the role of mineral binding in preserving the protein sequence in ostrich (Struthionidae) eggshell, including from the palaeontological sites of Laetoli (3.8 Ma) and Olduvai Gorge (1.3 Ma) in Tanzania. By tracking protein diagenesis back in time we find consistent patterns of preservation, demonstrating authenticity of the surviving sequences. Molecular dynamics simulations of struthiocalcin-1 and -2, the dominant proteins within the eggshell, reveal that distinct domains bind to the mineral surface. It is the domain with the strongest calculated binding energy to the calcite surface that is selectively preserved. Thermal age calculations demonstrate that the Laetoli and Olduvai peptides are 50 times older than any previously authenticated sequence (equivalent to ~16 Ma at a constant 10°C).

Research paper thumbnail of R. Dale Guthrie: Frozen Fauna of the Mammoth Steppe: The Story of Blue Babe

J Vertebrate Paleontol, 1990

Research paper thumbnail of Hunter-Gatherer Foraging Strategies in Tropical Grasslands: Model Building and Testing in the East African Middle and Later Stone Age

Journal of Anthropological Archaeology, Sep 1, 1997

Research paper thumbnail of Large mammal skeletal element transport: applying foraging theory in a complex taphonomic system

Journal of Taphonomy, 2003

Research paper thumbnail of Later quaternary extinct ungulates of East Africa and palaeoenvironmental implications