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Papers by Guy Hanley

12. Prodilis belinda Gordon and Hanley, new species Description. Male... more 12. Prodilis belinda Gordon and Hanley, new species Description. Male holotype. Length 2.3 mm, width 1.6 mm; body elongate oval, slender, elytron with side nearly straight, wider than pronotal base, widest at middle of elytra. Dorsal surface entirely shiny without trace of microsculpture. Color black; head black with yellow macula on frons, macula bisected by 2 longitudinal, narrow, black vittae (Fig. 382); pronotum black except lateral margin narrowly reddish brown; elytron with 2 yellow macula, anterior macula at humeral angle occupying most of anterior ½ of pronotum, large, irregularly square, posterior macula smaller, on apical portion of apical declivity (Fig. 380); antenna, legs, entire ventral surface except prosternum yellow; apical maxillary palpomere with apical ½ dark brown; prosternum yellowish red; abdomen yellow. Head punctures small, separated by a diameter or less; pronotal punctures as large as on head, separated by less than tw...

Faunistical description of ladybird assemblages from the canopies of agricultural plants and orch... more Faunistical description of ladybird assemblages from the canopies of agricultural plants and orchard trees in northeastern Iran and determination of the indicator ladybird species that are representative of the region are the main goals of this survey. In total, 21 coccinellid species belonging to 16 genera were collected and identified. One previously recorded species (Bulaea lichatschovii (Hummel, 1827)) was not confirmed. Global distribution, list of known host plants, and prey species are provided for each species, along with latitude, longitude, altitude, and sampling methodology. Multivariate analysis of ladybird species assemblages composition and environmental variables (canonical correspondence analysis, CCA) showed that most ladybird species analysed did not express any strong preference for the environmental variables mainly because of intercorrelated characteristics of sampling localities. Seasonality was clear in three species; Adalia decempunctata was the earliest spring species while Coccinella septempunctata and Vibidia duodecimguttata were abundant at the end of season. Although analysis of food webs showed that twenty one collected ladybird species were directly connected to 30 host plant species, this does not represent specific food relationships because prey species of predatory ladybirds were not sampled and identified. Instead, this may probably be interpreted as choice of microhabitats.

The Coleopterists Bulletin, 2020

BioOne Complete (complete.BioOne.org) is a full-text database of 200 subscribed and open-access t... more BioOne Complete (complete.BioOne.org) is a full-text database of 200 subscribed and open-access titles in the biological, ecological, and environmental sciences published by nonprofit societies, associations, museums, institutions, and presses.

Gordon, Robert D., Hanley, Guy A. (2014): Systematic revision of American Glaresidae (Coleoptera:... more Gordon, Robert D., Hanley, Guy A. (2014): Systematic revision of American Glaresidae (Coleoptera: Scarabaeoidea). Insecta Mundi 2014 (333): 1-91, DOI: http://doi.org/10.5281/zenodo.4645613

is a journal primarily devoted to insect systematics, but articles can be published on any nonmar... more is a journal primarily devoted to insect systematics, but articles can be published on any nonmarine arthropod. Topics considered for publication include systematics, taxonomy, nomenclature, checklists, faunal works, and natural history. Insecta Mundi will not consider works in the applied sciences (i.e. medical entomology, pest control research, etc.), and no longer publishes book reviews or editorials. Insecta Mundi publishes original research or discoveries in an inexpensive and timely manner, distributing them free via open access on the internet on the date of publication. Insecta Mundi is referenced or abstracted by several sources including the Zoological Record, CAB Abstracts, etc. Insecta Mundi is published irregularly throughout the year, with completed manuscripts assigned an individual number. Manuscripts must be peer reviewed prior to submission, after which they are reviewed by the editorial board to ensure quality. One author of each submitted manuscript must be a current member of the Center for Systematic Entomology. Manuscript preparation guidelines are availablr at the CSE website.

35. Neaporia felicia Gordon and Hanley, new species Description. Male... more 35. Neaporia felicia Gordon and Hanley, new species Description. Male holotype. Length 1.4 mm, width 1.0 mm; body elongate oval, elytron with side slightly rounded, wider than pronotal base, widest at middle of elytra. Dorsal surface shiny except elytron with trace of microsculpture. Color black; head black (Fig. 225); pronotum with reflexed lateral margin reddish brown; elytron reddish yellow with triangular, black basal macula extended from base at scutellum posteriorly to middle of elytron, apex of elytron narrowly brown (Fig. 224); antenna, epipleuron, legs yellow; mouthparts yellow except terminal maxillary palpomere dark brown with yellow ventral surface; ventral surface reddish yellow except abdomen yellowish brown. Head punctures small, separated by a diameter or less; pronotal punctures as large as on head, separated by less than twice a diameter; elytral punctures larger than on pronotum, separated by a diameter or less; prosternal pun...

15. Neaporia jenny Gordon and Hanley, new species Description. Male&l... more 15. Neaporia jenny Gordon and Hanley, new species Description. Male holotype. Length 1.6 mm, width 1.2 mm; body elongate oval, elytron with side weakly rounded, wider than pronotal base, widest anterior to middle of elytra. Dorsal surface entirely shiny, lacking microsculpture. Color brown (Fig. 129); head dark brown; pronotum with lateral 1/3 yellowish brown, not seen in image; mouthparts yellow except extreme apex of apical maxillary palpomere dark brown; antenna, mouthparts, legs yellow; apical 2 abdominal ventrites yellowish brown. Head punctures large, separated by less than a diameter; pronotal punctures about as large as on head, separated by less than a diameter; elytral punctures larger than on pronotum, separated by a diameter or less; prosternal, mesosternal punctures large, separated by less than a diameter; metasternum with large punctures along anterior and lateral borders, punctures on remaining surface small, sparse; punctures on...

7. Neaporia marsha Gordon and Hanley, new species Description. Male&l... more 7. Neaporia marsha Gordon and Hanley, new species Description. Male holotype. Length approximately 3.0 mm, width 2.2 mm; body short, wide, elytra much wider than pronotal base, widest at anterior 1/3 of elytra. Elytral surface entirely shiny, lacking microsculpture. Color of head and pronotum unknown, elytron pale red except black macula extended from base to apex medially, macula narrow on basal margin inside humeral callus, narrowly extended along sutural margin to just posterior to middle then widened to occupy remainder of elytron (Fig. 82); venter and legs yellow except prosternum slightly darker than remainder of venter. Elytral punctures separated by a diameter or less; prosternal punctures large, separated by less than a diameter; metasternal depression impunctate, lateral punctures large, separated by less than a diameter; punctures on abdominal ventrites 1, 2 large, separated by less than 3 times a diameter, inside of postcoxal lines l...

Figures 302-317. 302-306). Dilatitibialis ellen. 302) Habitus. 303) Phallobase ventral. 304) Phal... more Figures 302-317. 302-306). Dilatitibialis ellen. 302) Habitus. 303) Phallobase ventral. 304) Phallobase lateral. 305) Sipho. 306) Enlarged siphonal apex. 307-312) Dilatitibialis ceciliae. 307) Habitus. 308) Phallobase ventral. 309) Phallobase lateral. 310) Enlarged siphonal apex. 311) Sipho. 312) Female genitalia. 313-317) Dilatitibialis elaine. 313) Habitus. 314) Phallobase ventral. 315) Phallobase lateral. 316) Sipho. 317) Enlarged siphonal apex.

Figures 284-301. 284-289) Dilatitibialis shannon. 284) Habitus. 285) Phallobase ventral. 286) Pha... more Figures 284-301. 284-289) Dilatitibialis shannon. 284) Habitus. 285) Phallobase ventral. 286) Phallobase lateral. 287) Enlarged siphonal apex. 288) Sipho. 289) Female genitalia. 290-295) Dilatitibialis sheila. 290) Habitus. 291) Phallobase ventral. 292) Phallobase lateral. 293) Enlarged siphonal apex. 294) Sipho. 295) Female genitalia. 296- 301) Dilatitibialis ethel. 296) Habitus. 297) Phallobase ventral. 298) Phallobase lateral. 299) Enlarged siphonal apex. 300) Sipho. 301) Female genitalia.

Figures 318-325. 318-321) Dilatitibialis cruciferae. 318) Habitus. 319) Female genitalia. 320) Di... more Figures 318-325. 318-321) Dilatitibialis cruciferae. 318) Habitus. 319) Female genitalia. 320) Dilatitibialis gaynoni Habitus. 321) Female genitalia. 322-323) Dilatitibialis laterinotata. 322) Habitus. 323) Female genitalia. 324-325) Dilatitibialis marjorie. 324) Habitus. 325) Female genitalia.

Figures 268-283. 268-273) Dilatitibialis guttipennis. 268) Habitus. 269) Phallobase ventral. 270)... more Figures 268-283. 268-273) Dilatitibialis guttipennis. 268) Habitus. 269) Phallobase ventral. 270) Phallobase lateral. 271) Enlarged siphonal apex. 272) Sipho. 273) Female genitalia.274-278) Dilatitibialis josephine. 274) Habitus. 275) Phallobase ventral. 276) Phallobase lateral. 277) Sipho. 278) Enlarged siphonal apex. 279-283) Dilatitibialis thelma. 279) Habitus. 280) Phallobase ventral. 281) Phallobase lateral. 282) Sipho. 283) Enlarged siphonal apex.

Figures 235-250. 235-239) Dilatitibialis sherry. 235) Habitus. 236) Phallobase ventral. 237) Phal... more Figures 235-250. 235-239) Dilatitibialis sherry. 235) Habitus. 236) Phallobase ventral. 237) Phallobase lateral. 238) Sipho. 239) Enlarged siphonal apex. 240-245) Dilatitibialis cognata. 240) Habitus. 241) protibia. 242) Phallobase ventral. 243) Phallobase lateral. 244) Sipho. 245) Enlarged siphonal apex. 246-250) Dilatitibialis sylvia. 246) Habitus. 247) Phallobase ventral. 248) Phallobase lateral. 249) Sipho. 250) Enlarged siphonal apex.

Figures 368-385. Cyrea spp. 369-373) Cyrea dolores. 369) Habitus. 370) Phallobase ventral. 371) P... more Figures 368-385. Cyrea spp. 369-373) Cyrea dolores. 369) Habitus. 370) Phallobase ventral. 371) Phallobase lateral. 372) Sipho. 373) Female genitalia. 374-378) Cyrea erica. 374) Habitus. 375) Phallobase ventral. 376) Phallobase lateral. 377) Sipho. 378) Enlarged siphonal apex. 379-385) Cyrea tessulata. 379) Habitus. 380) Proleg. 381) Phallobase ventral. 382) Phallobase lateral. 383) Sipho. 384) Enlarged siphonal apex. 385) Female spermathecal capsule.

Figures 421-435. Cyrea spp. 421-425) Cyrea marion. 421) Habitus. 422) Phallobase ventral. 423) Ph... more Figures 421-435. Cyrea spp. 421-425) Cyrea marion. 421) Habitus. 422) Phallobase ventral. 423) Phallobase lateral. 424) Sipho. 425) Enlarged siphonal apex. 426-430) Cyrea dana. 426) Habitus. 427) Phallobase ventral. 428) Phallobase lateral. 429) Sipho. 430) Enlarged siphonal apex. 431-435) Cyrea stacy. 431) Habitus. 432) Phallobase ventral. 433) Phallobase lateral. 434) Sipho. 435) Enlarged siphonal apex.

Figures 572-589. Cyrea spp. 572-576) Cyrea georgia. 572) Habitus. 573) Phallobase ventral. 574) P... more Figures 572-589. Cyrea spp. 572-576) Cyrea georgia. 572) Habitus. 573) Phallobase ventral. 574) Phallobase lateral. 575) Sipho. 576) Enlarged siphonal apex. 577-583) Cyrea ruizi. 577) Habitus. 578) Proleg. 579) Phallobase ventral. 580) Phallobase lateral. 581) Sipho. 582) Enlarged siphonal apex. 583) Female genitalia. 584-589) Cyrea constance. 584) Habitus. 585) Phallobase ventral. 586) Phallobase lateral. 587) Sipho. 588) Enlarged siphonal apex. 589) Female spermathecal capsule.

Figures 554-571. Cyrea spp. 554-559) Cyrea tamara. 554) Habitus. 555) Phallobase ventral. 556) Ph... more Figures 554-571. Cyrea spp. 554-559) Cyrea tamara. 554) Habitus. 555) Phallobase ventral. 556) Phallobase lateral. 557) Sipho. 558) Enlarged siphonal apex. 559) Female genitalia. 560-565) Cyrea exclamationis. 560) Habitus. 561) Phallobase ventral. 562) Phallobase lateral. 563) Sipho. 564) Enlarged siphonal apex. 565) Female genitalia. 566-571) Cyrea joy. 566) Habitus. 567) Phallobase ventral. 568) Phallobase lateral. 569) Sipho. 570) Enlarged siphonal apex. 571) Female genitalia.

Figures 649-654. Cyrea spp. 649-650) Cyrea octupla. 649) Habitus. 650) Female genitalia. 651) Cyr... more Figures 649-654. Cyrea spp. 649-650) Cyrea octupla. 649) Habitus. 650) Female genitalia. 651) Cyrea flavocalceata. 651) Habitus. 652-654) Cyrea sexguttata. 652) Habitus. 653) Proleg. 654) Female genitalia.

The Coleopterists Bulletin, 2020