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Papers by Shivanthi Manickasingham

Research paper thumbnail of Qualitative and quantitative effects of CD28/B7-mediated costimulation on naive T cells in vitro

The Journal of Immunology, Oct 15, 1998

The CD28/B7 system provides costimulatory signals necessary for optimal T cell activation. We hav... more The CD28/B7 system provides costimulatory signals necessary for optimal T cell activation. We have examined the effects of blocking B7.1 and/or B7.2 in an in vitro system using TCR transgenic T cells specific for myelin basic protein. Activation of naive T cells was found to be B7.2 dependent and not dependent on the presence of B7.1 molecules. However, increasing the strength of signal through the TCR using peptide analogues with higher affinity for MHC compensated for blockade of B7.2 molecules, suggesting that signal 1 alone can be sufficient for the activation of naive T cells. The role of B7 molecules in the differentiation

Research paper thumbnail of Kinetics of Peptide Uptake and Tissue Distribution Following a Single Intranasal Dose of Peptide

We have previously used intranasal (i.n.) peptide application to induce mucosal tolerance in expe... more We have previously used intranasal (i.n.) peptide application to induce mucosal tolerance in experimental autoimmune encephalomyelitis (EAE). This strategy, however, appeared to give rise to similar phenomena of tolerance observed as a result of systemic administration of soluble antigenic peptide. We were interested, therefore, in the uptake and tissue distribution of peptide following i.n. treatment. In the H-2u mouse model of EAE, the highly tolerogenic peptide analogue Ac1-9[4Y] of myelin basic protein (MBP) displays high affinity binding to Au MHC class II. For the purpose of the present study this peptide was synthesised to contain a tritiated acetyl group and a protocol was developed to recover radioactivity in solubilised tissues taken at various times after [3H]Ac1-9[4Y] i.n.. Radiolabel loads of the lung and gastro-intestinal tract were initially high but declined rapidly. Radiolabel uptake by blood and lymphoid tissues followed similar kinetics with peak levels around 2.5-4 hours after i.n. administration. Concentrations were high in the draining cervical lymph nodes (CLN) but also reached significant levels in the spleen and 'nondraining' inguinal lymph nodes. The presence of intact antigenic peptide was demonstrated in spleens and CLN from Ac1-9[4Y] i.n. treated mice. Cell suspensions prepared from these tissues at selected time points after peptide i.n. were able to stimulate peptide-specific T cell lines up to at least one day after peptide i.n., suggesting long lasting formation of stable Au-Ac1-9[4Y] complexes in vivo.

Research paper thumbnail of Microbial and T Cell-Derived Stimuli Regulate Antigen Presentation by Dendritic Cells In Vivo

B cells and dendritic cells (DC) internalize and degrade exogenous Ags and present them as peptid... more B cells and dendritic cells (DC) internalize and degrade exogenous Ags and present them as peptides bound to MHC class II molecules for scrutiny by CD4(+) T cells. Here we use an Ab specific for a processed form of the model Ag, hen egg lysozyme (HEL), to demonstrate that this protein is not efficiently presented by lymph node DC following s.c. immunization. HEL presentation by the DC can be dramatically enhanced upon coinjection of a microbial adjuvant, which appears to act by enhancing peptide loading onto MHC class II. CD40 cross-linking or the presence of a high frequency of T cells specific for HEL can similarly improve presentation by DC in vivo. For any of these activating stimuli, CD8alpha(+) DC consistently display the highest proportion of HEL-loaded MHC class II molecules. These data indicate that exogenous Ags can be displayed to T cells in lymphoid tissues by a large cohort of resident DC whose presentation is regulated by innate and adaptive stimuli. Our data further reveal the existence of a feedback mechanism that augments Ag presentation during cognate APC-T cell interactions.

Research paper thumbnail of Fine specificity of the myelin-reactive T cell repertoire: implications for TCR antagonism in autoimmunity

Journal of immunology (Baltimore, Md. : 1950), 1998

The use of altered peptide ligands (APL) to modulate T cell responses has been suggested as a mea... more The use of altered peptide ligands (APL) to modulate T cell responses has been suggested as a means of treating T cell-mediated autoimmune disorders. We have assessed the therapeutic potential of TCR antagonist peptides in autoimmunity using murine experimental autoimmune encephalomyelitis (EAE) as a model. The Tg4 transgenic mouse expresses an MHC class II-restricted TCR specific for the immunodominant encephalitogenic epitope of myelin basic protein, Ac1-9 (acetylated N-terminal nonamer). We have used T cell lines derived from Tg4 mice to define the TCR contact residues within Ac1-9. APL with appropriate substitutions at the primary TCR contact residue were effective antagonists of Tg4 T cells. These antagonist APL, however, were found to induce EAE in susceptible, nontransgenic strains of mice. Underlying this, the Ac1-9-specific T cell repertoire of normal mice, rather than reflecting the Tg4 phenotype, showed considerable diversity in fine specificity and was able to respond to...

Research paper thumbnail of Triggering Dendritic Cell Subsets to CD40 the Cytokine Response of Murine Independent Pathways Determines Receptor-Dependent and Microbial Recognition Via Toll-Like

Research paper thumbnail of TCR Antagonism in Autoimmunity T Cell Repertoire: Implications for Fine Specificity of the Myelin-Reactive

Research paper thumbnail of Qualitative and Quantitative Effects of CD28/B7-Mediated Costimulation on Naive T Cells In Vitro1

regulation of IL-2 and IFN-g production by T cells primed in the presence of anti-B7.2 mAb was pa... more regulation of IL-2 and IFN-g production by T cells primed in the presence of anti-B7.2 mAb was partially overcome when high affinity peptide analogues were used to restimulate T cells. In contrast, a significant down-regulation of the differentiation of cells producing Th-2 cytokines was observed in the presence of anti-B7 Abs. Differentiation of IL-4-secreting cells was influenced by both B7.1

Research paper thumbnail of Microbial Recognition Via Toll-Like Receptor-Dependent and -Independent Pathways Determines the Cytokine Response of Murine Dendritic Cell Subsets to CD40 Triggering

The Journal of Immunology, 2002

Research paper thumbnail of Fine specificity of myelin basic protein specific T cells: Implications for TCR antagonism

Immunology Letters, 1997

the T peripheral blood cells response to I? aeruginosa antigens and the nature of these ligands f... more the T peripheral blood cells response to I? aeruginosa antigens and the nature of these ligands for ab and y6 T lymphocytes.

Research paper thumbnail of Mature T cell seeks antigen for meaningful relationship in lymph node

Research paper thumbnail of Kinetics of Peptide Uptake and Tissue Distribution Following a Single Intranasal Dose of Peptide

Immunological Investigations, 2000

We have previously used intranasal (i.n.) peptide application to induce mucosal tolerance in expe... more We have previously used intranasal (i.n.) peptide application to induce mucosal tolerance in experimental autoimmune encephalomyelitis (EAE). This strategy, however, appeared to give rise to similar phenomena of tolerance observed as a result of systemic administration of soluble antigenic peptide. We were interested, therefore, in the uptake and tissue distribution of peptide following i.n. treatment. In the H-2u mouse model of EAE, the highly tolerogenic peptide analogue Ac1-9[4Y] of myelin basic protein (MBP) displays high affinity binding to Au MHC class II. For the purpose of the present study this peptide was synthesised to contain a tritiated acetyl group and a protocol was developed to recover radioactivity in solubilised tissues taken at various times after [3H]Ac1-9[4Y] i.n.. Radiolabel loads of the lung and gastro-intestinal tract were initially high but declined rapidly. Radiolabel uptake by blood and lymphoid tissues followed similar kinetics with peak levels around 2.5-4 hours after i.n. administration. Concentrations were high in the draining cervical lymph nodes (CLN) but also reached significant levels in the spleen and 'nondraining' inguinal lymph nodes. The presence of intact antigenic peptide was demonstrated in spleens and CLN from Ac1-9[4Y] i.n. treated mice. Cell suspensions prepared from these tissues at selected time points after peptide i.n. were able to stimulate peptide-specific T cell lines up to at least one day after peptide i.n., suggesting long lasting formation of stable Au-Ac1-9[4Y] complexes in vivo.

Research paper thumbnail of Conditioning of Dendritic Cells by Pathogen-Derived Stimuli

Immunobiology, Jan 1, 2001

AD Edwards SP Manickasingham R. Spörri SS Diebold O. Schulz A. Sher T. Kaisho S. Akira C. Reise S... more AD Edwards SP Manickasingham R. Spörri SS Diebold O. Schulz A. Sher T. Kaisho S. Akira C. Reise Sousa 2001. Microbial recognition via toll-like receptor-dependent and -independent pathways determines the cytokine response of murine dendritic cell subsets to CD40 triggering. Submitted. ... Copyright © 2001 Urban & Fischer Verlag. Published by Elsevier GmbH All rights reserved.

Research paper thumbnail of The Ability of Murine Dendritic Cell Subsets to Direct T Helper Cell Differentiation is Dependent on Microbial Signals

European journal …, Jan 1, 2003

Dendritic cells (DC) initiate T cell responses and direct the class of T cell immunity through th... more Dendritic cells (DC) initiate T cell responses and direct the class of T cell immunity through the production of Th-polarizing cytokines. In the mouse, immunization with CD8alpha(+) DC has led to Th1 priming whereas immunization with CD8alpha(-) DC has been associated with Th2 induction. Here, we use a direct T cell priming assay in vitro to re-examine the Th-directing potential of total DC or purified CD4(+) DC, CD8alpha(+) DC or CD4(-) CD8alpha(-) (double-negative; DN) DC subsets from mouse spleen. We show that the default Th effector phenotype induced by priming with DC depends on the protocol used for T cell purification, the T cell:antigen-presenting cell ratio and the antigen dose but is only marginally affected by DC subtype. All DC subsets can direct increased Th1 development in response to microbial stimuli known to elicit IL-12 production. Similarly, all subsets can suppress Th1 development and allow Th2 cellsto expand upon exposure to IL-10-inducing microbial agents. The flexibility of DC in directing Th development in function of microbial signals argues against the notion of pre-determined "DC1" and "DC2" subsets and suggests that multiple DC subtypes can direct an appropriate Th response to different classes of infectious agents.

Research paper thumbnail of CELLULAR IMMUNOLOGY AND IMMUNE REGULATION-Microbial Recognition Via Toll-Uke Receptor-Dependent and-Independent Pathways Determines the Cytokine Response of Murine Dendritic Cell Subsets to CD40

Journal of Immunology, 2002

Research paper thumbnail of CD40 Triggering of Heterodimeric IL-12 P70 Production by Dendritic Cells In Vivo Requires a Microbial Priming Signal

Research paper thumbnail of Qualitative and quantitative effects of CD28/B7-mediated costimulation on naive T cells in vitro

The Journal of Immunology, Oct 15, 1998

The CD28/B7 system provides costimulatory signals necessary for optimal T cell activation. We hav... more The CD28/B7 system provides costimulatory signals necessary for optimal T cell activation. We have examined the effects of blocking B7.1 and/or B7.2 in an in vitro system using TCR transgenic T cells specific for myelin basic protein. Activation of naive T cells was found to be B7.2 dependent and not dependent on the presence of B7.1 molecules. However, increasing the strength of signal through the TCR using peptide analogues with higher affinity for MHC compensated for blockade of B7.2 molecules, suggesting that signal 1 alone can be sufficient for the activation of naive T cells. The role of B7 molecules in the differentiation

Research paper thumbnail of Kinetics of Peptide Uptake and Tissue Distribution Following a Single Intranasal Dose of Peptide

We have previously used intranasal (i.n.) peptide application to induce mucosal tolerance in expe... more We have previously used intranasal (i.n.) peptide application to induce mucosal tolerance in experimental autoimmune encephalomyelitis (EAE). This strategy, however, appeared to give rise to similar phenomena of tolerance observed as a result of systemic administration of soluble antigenic peptide. We were interested, therefore, in the uptake and tissue distribution of peptide following i.n. treatment. In the H-2u mouse model of EAE, the highly tolerogenic peptide analogue Ac1-9[4Y] of myelin basic protein (MBP) displays high affinity binding to Au MHC class II. For the purpose of the present study this peptide was synthesised to contain a tritiated acetyl group and a protocol was developed to recover radioactivity in solubilised tissues taken at various times after [3H]Ac1-9[4Y] i.n.. Radiolabel loads of the lung and gastro-intestinal tract were initially high but declined rapidly. Radiolabel uptake by blood and lymphoid tissues followed similar kinetics with peak levels around 2.5-4 hours after i.n. administration. Concentrations were high in the draining cervical lymph nodes (CLN) but also reached significant levels in the spleen and 'nondraining' inguinal lymph nodes. The presence of intact antigenic peptide was demonstrated in spleens and CLN from Ac1-9[4Y] i.n. treated mice. Cell suspensions prepared from these tissues at selected time points after peptide i.n. were able to stimulate peptide-specific T cell lines up to at least one day after peptide i.n., suggesting long lasting formation of stable Au-Ac1-9[4Y] complexes in vivo.

Research paper thumbnail of Microbial and T Cell-Derived Stimuli Regulate Antigen Presentation by Dendritic Cells In Vivo

B cells and dendritic cells (DC) internalize and degrade exogenous Ags and present them as peptid... more B cells and dendritic cells (DC) internalize and degrade exogenous Ags and present them as peptides bound to MHC class II molecules for scrutiny by CD4(+) T cells. Here we use an Ab specific for a processed form of the model Ag, hen egg lysozyme (HEL), to demonstrate that this protein is not efficiently presented by lymph node DC following s.c. immunization. HEL presentation by the DC can be dramatically enhanced upon coinjection of a microbial adjuvant, which appears to act by enhancing peptide loading onto MHC class II. CD40 cross-linking or the presence of a high frequency of T cells specific for HEL can similarly improve presentation by DC in vivo. For any of these activating stimuli, CD8alpha(+) DC consistently display the highest proportion of HEL-loaded MHC class II molecules. These data indicate that exogenous Ags can be displayed to T cells in lymphoid tissues by a large cohort of resident DC whose presentation is regulated by innate and adaptive stimuli. Our data further reveal the existence of a feedback mechanism that augments Ag presentation during cognate APC-T cell interactions.

Research paper thumbnail of Fine specificity of the myelin-reactive T cell repertoire: implications for TCR antagonism in autoimmunity

Journal of immunology (Baltimore, Md. : 1950), 1998

The use of altered peptide ligands (APL) to modulate T cell responses has been suggested as a mea... more The use of altered peptide ligands (APL) to modulate T cell responses has been suggested as a means of treating T cell-mediated autoimmune disorders. We have assessed the therapeutic potential of TCR antagonist peptides in autoimmunity using murine experimental autoimmune encephalomyelitis (EAE) as a model. The Tg4 transgenic mouse expresses an MHC class II-restricted TCR specific for the immunodominant encephalitogenic epitope of myelin basic protein, Ac1-9 (acetylated N-terminal nonamer). We have used T cell lines derived from Tg4 mice to define the TCR contact residues within Ac1-9. APL with appropriate substitutions at the primary TCR contact residue were effective antagonists of Tg4 T cells. These antagonist APL, however, were found to induce EAE in susceptible, nontransgenic strains of mice. Underlying this, the Ac1-9-specific T cell repertoire of normal mice, rather than reflecting the Tg4 phenotype, showed considerable diversity in fine specificity and was able to respond to...

Research paper thumbnail of Triggering Dendritic Cell Subsets to CD40 the Cytokine Response of Murine Independent Pathways Determines Receptor-Dependent and Microbial Recognition Via Toll-Like

Research paper thumbnail of TCR Antagonism in Autoimmunity T Cell Repertoire: Implications for Fine Specificity of the Myelin-Reactive

Research paper thumbnail of Qualitative and Quantitative Effects of CD28/B7-Mediated Costimulation on Naive T Cells In Vitro1

regulation of IL-2 and IFN-g production by T cells primed in the presence of anti-B7.2 mAb was pa... more regulation of IL-2 and IFN-g production by T cells primed in the presence of anti-B7.2 mAb was partially overcome when high affinity peptide analogues were used to restimulate T cells. In contrast, a significant down-regulation of the differentiation of cells producing Th-2 cytokines was observed in the presence of anti-B7 Abs. Differentiation of IL-4-secreting cells was influenced by both B7.1

Research paper thumbnail of Microbial Recognition Via Toll-Like Receptor-Dependent and -Independent Pathways Determines the Cytokine Response of Murine Dendritic Cell Subsets to CD40 Triggering

The Journal of Immunology, 2002

Research paper thumbnail of Fine specificity of myelin basic protein specific T cells: Implications for TCR antagonism

Immunology Letters, 1997

the T peripheral blood cells response to I? aeruginosa antigens and the nature of these ligands f... more the T peripheral blood cells response to I? aeruginosa antigens and the nature of these ligands for ab and y6 T lymphocytes.

Research paper thumbnail of Mature T cell seeks antigen for meaningful relationship in lymph node

Research paper thumbnail of Kinetics of Peptide Uptake and Tissue Distribution Following a Single Intranasal Dose of Peptide

Immunological Investigations, 2000

We have previously used intranasal (i.n.) peptide application to induce mucosal tolerance in expe... more We have previously used intranasal (i.n.) peptide application to induce mucosal tolerance in experimental autoimmune encephalomyelitis (EAE). This strategy, however, appeared to give rise to similar phenomena of tolerance observed as a result of systemic administration of soluble antigenic peptide. We were interested, therefore, in the uptake and tissue distribution of peptide following i.n. treatment. In the H-2u mouse model of EAE, the highly tolerogenic peptide analogue Ac1-9[4Y] of myelin basic protein (MBP) displays high affinity binding to Au MHC class II. For the purpose of the present study this peptide was synthesised to contain a tritiated acetyl group and a protocol was developed to recover radioactivity in solubilised tissues taken at various times after [3H]Ac1-9[4Y] i.n.. Radiolabel loads of the lung and gastro-intestinal tract were initially high but declined rapidly. Radiolabel uptake by blood and lymphoid tissues followed similar kinetics with peak levels around 2.5-4 hours after i.n. administration. Concentrations were high in the draining cervical lymph nodes (CLN) but also reached significant levels in the spleen and 'nondraining' inguinal lymph nodes. The presence of intact antigenic peptide was demonstrated in spleens and CLN from Ac1-9[4Y] i.n. treated mice. Cell suspensions prepared from these tissues at selected time points after peptide i.n. were able to stimulate peptide-specific T cell lines up to at least one day after peptide i.n., suggesting long lasting formation of stable Au-Ac1-9[4Y] complexes in vivo.

Research paper thumbnail of Conditioning of Dendritic Cells by Pathogen-Derived Stimuli

Immunobiology, Jan 1, 2001

AD Edwards SP Manickasingham R. Spörri SS Diebold O. Schulz A. Sher T. Kaisho S. Akira C. Reise S... more AD Edwards SP Manickasingham R. Spörri SS Diebold O. Schulz A. Sher T. Kaisho S. Akira C. Reise Sousa 2001. Microbial recognition via toll-like receptor-dependent and -independent pathways determines the cytokine response of murine dendritic cell subsets to CD40 triggering. Submitted. ... Copyright © 2001 Urban & Fischer Verlag. Published by Elsevier GmbH All rights reserved.

Research paper thumbnail of The Ability of Murine Dendritic Cell Subsets to Direct T Helper Cell Differentiation is Dependent on Microbial Signals

European journal …, Jan 1, 2003

Dendritic cells (DC) initiate T cell responses and direct the class of T cell immunity through th... more Dendritic cells (DC) initiate T cell responses and direct the class of T cell immunity through the production of Th-polarizing cytokines. In the mouse, immunization with CD8alpha(+) DC has led to Th1 priming whereas immunization with CD8alpha(-) DC has been associated with Th2 induction. Here, we use a direct T cell priming assay in vitro to re-examine the Th-directing potential of total DC or purified CD4(+) DC, CD8alpha(+) DC or CD4(-) CD8alpha(-) (double-negative; DN) DC subsets from mouse spleen. We show that the default Th effector phenotype induced by priming with DC depends on the protocol used for T cell purification, the T cell:antigen-presenting cell ratio and the antigen dose but is only marginally affected by DC subtype. All DC subsets can direct increased Th1 development in response to microbial stimuli known to elicit IL-12 production. Similarly, all subsets can suppress Th1 development and allow Th2 cellsto expand upon exposure to IL-10-inducing microbial agents. The flexibility of DC in directing Th development in function of microbial signals argues against the notion of pre-determined "DC1" and "DC2" subsets and suggests that multiple DC subtypes can direct an appropriate Th response to different classes of infectious agents.

Research paper thumbnail of CELLULAR IMMUNOLOGY AND IMMUNE REGULATION-Microbial Recognition Via Toll-Uke Receptor-Dependent and-Independent Pathways Determines the Cytokine Response of Murine Dendritic Cell Subsets to CD40

Journal of Immunology, 2002

Research paper thumbnail of CD40 Triggering of Heterodimeric IL-12 P70 Production by Dendritic Cells In Vivo Requires a Microbial Priming Signal