Spase Shumka - Academia.edu (original) (raw)
Papers by Spase Shumka
Its well known that eutrophication processes caused by an enrichment in nutrients results in an i... more Its well known that eutrophication processes caused by an enrichment in nutrients results in an increase of photosynthetic biomass. In the water ecology however, the factors which drive the shift in species composition of the assemblage along trophic gradients are yet poorly understood (Reynolds, 1996).
... S. Grazhdani (B) Agricultural University of Tirana, Tirana, Albania e-mail: Spiro.grazhdani@ ... more ... S. Grazhdani (B) Agricultural University of Tirana, Tirana, Albania e-mail: Spiro.grazhdani@ yahoo.com 117 H. Gökçekus et al. ... Further, small organic particles such as phytoplankton and detritus, can be ingested and passed along to higher organisms. ...
Desalination, 2007
The combination of the mountainous terrain of Albania and a seasonally wet Mediterranean climate ... more The combination of the mountainous terrain of Albania and a seasonally wet Mediterranean climate has led to some of the most extreme erosion in Europe. New methodology is used in this work to estimate soil erosion risk and provide a framework for soil conservation that can effectively and economically control soil loss. A onedimensional hydrology, vegetation and erosion model (RDI/CSEP) is used to calculate accumulative soil erosion. This model provides an integrating the climatic, topography and associated vegetation components into soil erosion modeling. The forecast runoff, accumulated across the frequency distribution of storms, is used to give a climatic potential, which is then appropriately combined with measures of topography and soil erodibility to estimate the expected rate of soil erosion at a resolution of 1 km.
Survival and Sustainability, 2010
... S. Grazhdani (B) Agricultural University of Tirana, Tirana, Albania e-mail: Spiro.grazhdani@ ... more ... S. Grazhdani (B) Agricultural University of Tirana, Tirana, Albania e-mail: Spiro.grazhdani@ yahoo.com 117 H. Gökçekus et al. ... Further, small organic particles such as phytoplankton and detritus, can be ingested and passed along to higher organisms. ...
Endangered Species Research, 2014
The subject of this paper is the water life and energy issue of concerns within Rivers of Vjosa, ... more The subject of this paper is the water life and energy issue of concerns within Rivers of Vjosa, Semani and Drini the main contributor with freshwater to south eastern Adriatic. The rivers of Albania are mostly natural; they have hardly been dammed or channeled, except for some existing dams in the middle part of the river courses. Therefore these rivers,
The Water Framework Directive of the EU provides the ground leading guidelines for meeting the st... more The Water Framework Directive of the EU provides the ground leading guidelines for meeting the standards in both water quality and quantity. This is affecting not only the member states, but also the countries that are in the process of integration. To that fact, based on possible scheme of monitoring of surface water, the microbiological aspects comprises a target to
On the Balkans thirteen river basins are transboundary, i.e. shared between two or more countries... more On the Balkans thirteen river basins are transboundary, i.e. shared between two or more countries. This illustrates the high relevance of developing appropriate management tools and policy strategies tailored to address transboundary water resource issues in this region. The DRIMON Project (www.drimon.net)
Water Resources Management, 2000
A limnological survey of a man-made reservoir in the central part of Albania was conducted from M... more A limnological survey of a man-made reservoir in the central part of Albania was conducted from May 2006 to May 2008. This water body is being constructed 15 years ago for water supply of the Tirana, capital city of Albania. The physical and chemical quality of water in this reservoir was determined and compared against universally accepted water quality standards. Water quality parameters relevant to the study area are discussed considering the ecosystem function and interrelationship among physical and chemical parameters with the biological ones. The water quality of traditional and modern reservoir was not significantly different except for a bed odour that occurs throughout of the summer. Due to the reservoirs life the eutrophication in the nowadays is not a major problem in the reservoir, but due to an intensive sedimentation rate it is predicted that will seriously affect the water body in the near future. Management strategies to improve the water use efficiency with particular reference to eutrophication are proposed. On the other hand it is recommended that an integrated approach following the IRBM criteria will be the best way of management and administration.
A Multidisciplinary Approach, 2011
... 2 Priscoli, JD and Wolf, AT (2009) Managing and Transforming Water Conflicts, Cambridge Unive... more ... 2 Priscoli, JD and Wolf, AT (2009) Managing and Transforming Water Conflicts, Cambridge University ... embedded in local institutions and practices, providing a useful source of knowledge ... New economic zones were established for intensive rice cultivation: typically three crops ...
Environmental Monitoring and Assessment, 2000
Bovilla Lake is a reservoir constructed 12 years ago for supplying the city of Tirana (Albania) m... more Bovilla Lake is a reservoir constructed 12 years ago for supplying the city of Tirana (Albania) mainly with drinking water. It has a surface area of 4.6 km 2 , a maximum depth of originally 60 m and is monomictic with a stratification Electronic supplementary material The online version of this article (
Genetics, selection, evolution : GSE, 2009
To determine the genetic architecture of trout in Albania, 87 individuals were collected from 19 ... more To determine the genetic architecture of trout in Albania, 87 individuals were collected from 19 riverine and lacustrine sites in Albania, FYROM and Greece. All individuals were analyzed for sequence variation in the mtDNA control region. Among fourteen haplotypes detected, four previously unpublished haplotypes, bearing a close relationship to haplotypes of the Adriatic and marmoratus lineages of Salmo trutta, were revealed. Ten previously described haplotypes, characteristic of S. ohridanus, S. letnica and the Adriatic and Mediterranean lineages of S. trutta, were also detected. Haplotypes detected in this study were placed in a well supported branch of S. ohridanus, and a cluster of Mediterranean -Adriatic -marmoratus haplotypes, which were further delimited into three subdivisions of Mediterranean, marmoratus, and a previously non-described formation of four Adriatic haplotypes (Balkan cluster). Haplotypes of the Balkan cluster and the other Adriatic haplotypes, do not represent a contiguous haplotype lineage and appear not to be closely related, indicating independent arrivals into the Adriatic drainage and suggesting successive colonization events. Despite the presence of marmoratus haplotypes in Albania, no marbled phenotype was found, confirming previously reported findings that there is no association between this phenotype and marmoratus haplotypes.
Molecular phylogenetics and evolution, 2010
Freshwater fauna of ancient lakes frequently contain endemic taxa thought to have originated duri... more Freshwater fauna of ancient lakes frequently contain endemic taxa thought to have originated during the long existence of these lakes, yet uncertainties remain as to whether they represent distinct genetic lineages with respect to more widespread relatives and to the relative roles of isolation and dispersal in their evolution. Phylogenetic analyses of sequence variation at nuclear and mitochondrial genes were used to examine these issues for the freshwater fish genus Barbus in two European ancient lake systems on the Balkan Peninsula. The nuclear and mitochondrial data yielded concordant phylogeographic patterns though incomplete sorting of nuclear haplotypes between some mitochondrial clades was detected. The distributions of two currently recognized species investigated here do not match the distributions of evolutionary lineages revealed by phylogenetic analyses. The Prespa barbel, Barbus prespensis, is not endemic to the lakes Prespa as previously thought but is instead found to be widespread in the southeastern Adriatic Sea basin, with a distribution largely corresponding to the basin of the now extinct Lake Maliq historically connected with Lake Prespa. On the other hand, a cryptic phylogenetic subdivision in a widespread species, B. rebeli, was discovered to be more distant from B. rebeli than from other Barbus species and to be endemic to the system of connected lakes Ohrid and Shkodra. The division coincides with the hydrogeographical boundary delimiting distributions of other freshwater fishes, and we suggest that this newly discovered evolutionary lineage represents a distinct species. These findings support the emerging pattern that endemic taxa have evolved not through isolation of individual lakes, but in systems of currently and historically interconnected lakes and their wider basins.
Molecular Phylogenetics and Evolution - MOL PHYLOGENET EVOL, 2010
Freshwater fauna of ancient lakes frequently contain endemic taxa thought to have originated duri... more Freshwater fauna of ancient lakes frequently contain endemic taxa thought to have originated during the long existence of these lakes, yet uncertainties remain as to whether they represent distinct genetic lineages with respect to more widespread relatives and to the relative roles of isolation and dispersal in their evolution. Phylogenetic analyses of sequence variation at nuclear and mitochondrial genes were used to examine these issues for the freshwater fish genus Barbus in two European ancient lake systems on the Balkan Peninsula. The nuclear and mitochondrial data yielded concordant phylogeographic patterns though incomplete sorting of nuclear haplotypes between some mitochondrial clades was detected. The distributions of two currently recognized species investigated here do not match the distributions of evolutionary lineages revealed by phylogenetic analyses. The Prespa barbel, Barbus prespensis, is not endemic to the lakes Prespa as previously thought but is instead found to be widespread in the southeastern Adriatic Sea basin, with a distribution largely corresponding to the basin of the now extinct Lake Maliq historically connected with Lake Prespa. On the other hand, a cryptic phylogenetic subdivision in a widespread species, B. rebeli, was discovered to be more distant from B. rebeli than from other Barbus species and to be endemic to the system of connected lakes Ohrid and Shkodra. The division coincides with the hydrogeographical boundary delimiting distributions of other freshwater fishes, and we suggest that this newly discovered evolutionary lineage represents a distinct species. These findings support the emerging pattern that endemic taxa have evolved not through isolation of individual lakes, but in systems of currently and historically interconnected lakes and their wider basins.
Genetics Selection Evolution, 2009
To determine the genetic architecture of trout in Albania, 87 individuals were collected from 19 ... more To determine the genetic architecture of trout in Albania, 87 individuals were collected from 19 riverine and lacustrine sites in Albania, FYROM and Greece. All individuals were analyzed for sequence variation in the mtDNA control region. Among fourteen haplotypes detected, four previously unpublished haplotypes, bearing a close relationship to haplotypes of the Adriatic and marmoratus lineages of Salmo trutta, were revealed. Ten previously described haplotypes, characteristic of S. ohridanus, S. letnica and the Adriatic and Mediterranean lineages of S. trutta, were also detected. Haplotypes detected in this study were placed in a well supported branch of S. ohridanus, and a cluster of Mediterranean -Adriatic -marmoratus haplotypes, which were further delimited into three subdivisions of Mediterranean, marmoratus, and a previously non-described formation of four Adriatic haplotypes (Balkan cluster). Haplotypes of the Balkan cluster and the other Adriatic haplotypes, do not represent a contiguous haplotype lineage and appear not to be closely related, indicating independent arrivals into the Adriatic drainage and suggesting successive colonization events. Despite the presence of marmoratus haplotypes in Albania, no marbled phenotype was found, confirming previously reported findings that there is no association between this phenotype and marmoratus haplotypes.
Melovski, D., Ivanov, Gj., Stojanov, A., Avukatov, V., Trajçe, A., Hoxha, B., von Arx, M., Breite... more Melovski, D., Ivanov, Gj., Stojanov, A., Avukatov, V., Trajçe, A., Hoxha, B., von Arx, M., Breitenmoser-Würsten, Ch., Hristovski, S., Shumka, S., Breitenmoser, U. (2013). Distribution and conservation status of the Balkan lynx (Lynx lynx balcanicus . Proceedings of the 4 th Congress of Ecologists of Macedonia with International Participation, Ohrid, 12-15 October 2012. Macedonian Ecological Society, Special issue 28, Skopje.
Its well known that eutrophication processes caused by an enrichment in nutrients results in an i... more Its well known that eutrophication processes caused by an enrichment in nutrients results in an increase of photosynthetic biomass. In the water ecology however, the factors which drive the shift in species composition of the assemblage along trophic gradients are yet poorly understood (Reynolds, 1996).
... S. Grazhdani (B) Agricultural University of Tirana, Tirana, Albania e-mail: Spiro.grazhdani@ ... more ... S. Grazhdani (B) Agricultural University of Tirana, Tirana, Albania e-mail: Spiro.grazhdani@ yahoo.com 117 H. Gökçekus et al. ... Further, small organic particles such as phytoplankton and detritus, can be ingested and passed along to higher organisms. ...
Desalination, 2007
The combination of the mountainous terrain of Albania and a seasonally wet Mediterranean climate ... more The combination of the mountainous terrain of Albania and a seasonally wet Mediterranean climate has led to some of the most extreme erosion in Europe. New methodology is used in this work to estimate soil erosion risk and provide a framework for soil conservation that can effectively and economically control soil loss. A onedimensional hydrology, vegetation and erosion model (RDI/CSEP) is used to calculate accumulative soil erosion. This model provides an integrating the climatic, topography and associated vegetation components into soil erosion modeling. The forecast runoff, accumulated across the frequency distribution of storms, is used to give a climatic potential, which is then appropriately combined with measures of topography and soil erodibility to estimate the expected rate of soil erosion at a resolution of 1 km.
Survival and Sustainability, 2010
... S. Grazhdani (B) Agricultural University of Tirana, Tirana, Albania e-mail: Spiro.grazhdani@ ... more ... S. Grazhdani (B) Agricultural University of Tirana, Tirana, Albania e-mail: Spiro.grazhdani@ yahoo.com 117 H. Gökçekus et al. ... Further, small organic particles such as phytoplankton and detritus, can be ingested and passed along to higher organisms. ...
Endangered Species Research, 2014
The subject of this paper is the water life and energy issue of concerns within Rivers of Vjosa, ... more The subject of this paper is the water life and energy issue of concerns within Rivers of Vjosa, Semani and Drini the main contributor with freshwater to south eastern Adriatic. The rivers of Albania are mostly natural; they have hardly been dammed or channeled, except for some existing dams in the middle part of the river courses. Therefore these rivers,
The Water Framework Directive of the EU provides the ground leading guidelines for meeting the st... more The Water Framework Directive of the EU provides the ground leading guidelines for meeting the standards in both water quality and quantity. This is affecting not only the member states, but also the countries that are in the process of integration. To that fact, based on possible scheme of monitoring of surface water, the microbiological aspects comprises a target to
On the Balkans thirteen river basins are transboundary, i.e. shared between two or more countries... more On the Balkans thirteen river basins are transboundary, i.e. shared between two or more countries. This illustrates the high relevance of developing appropriate management tools and policy strategies tailored to address transboundary water resource issues in this region. The DRIMON Project (www.drimon.net)
Water Resources Management, 2000
A limnological survey of a man-made reservoir in the central part of Albania was conducted from M... more A limnological survey of a man-made reservoir in the central part of Albania was conducted from May 2006 to May 2008. This water body is being constructed 15 years ago for water supply of the Tirana, capital city of Albania. The physical and chemical quality of water in this reservoir was determined and compared against universally accepted water quality standards. Water quality parameters relevant to the study area are discussed considering the ecosystem function and interrelationship among physical and chemical parameters with the biological ones. The water quality of traditional and modern reservoir was not significantly different except for a bed odour that occurs throughout of the summer. Due to the reservoirs life the eutrophication in the nowadays is not a major problem in the reservoir, but due to an intensive sedimentation rate it is predicted that will seriously affect the water body in the near future. Management strategies to improve the water use efficiency with particular reference to eutrophication are proposed. On the other hand it is recommended that an integrated approach following the IRBM criteria will be the best way of management and administration.
A Multidisciplinary Approach, 2011
... 2 Priscoli, JD and Wolf, AT (2009) Managing and Transforming Water Conflicts, Cambridge Unive... more ... 2 Priscoli, JD and Wolf, AT (2009) Managing and Transforming Water Conflicts, Cambridge University ... embedded in local institutions and practices, providing a useful source of knowledge ... New economic zones were established for intensive rice cultivation: typically three crops ...
Environmental Monitoring and Assessment, 2000
Bovilla Lake is a reservoir constructed 12 years ago for supplying the city of Tirana (Albania) m... more Bovilla Lake is a reservoir constructed 12 years ago for supplying the city of Tirana (Albania) mainly with drinking water. It has a surface area of 4.6 km 2 , a maximum depth of originally 60 m and is monomictic with a stratification Electronic supplementary material The online version of this article (
Genetics, selection, evolution : GSE, 2009
To determine the genetic architecture of trout in Albania, 87 individuals were collected from 19 ... more To determine the genetic architecture of trout in Albania, 87 individuals were collected from 19 riverine and lacustrine sites in Albania, FYROM and Greece. All individuals were analyzed for sequence variation in the mtDNA control region. Among fourteen haplotypes detected, four previously unpublished haplotypes, bearing a close relationship to haplotypes of the Adriatic and marmoratus lineages of Salmo trutta, were revealed. Ten previously described haplotypes, characteristic of S. ohridanus, S. letnica and the Adriatic and Mediterranean lineages of S. trutta, were also detected. Haplotypes detected in this study were placed in a well supported branch of S. ohridanus, and a cluster of Mediterranean -Adriatic -marmoratus haplotypes, which were further delimited into three subdivisions of Mediterranean, marmoratus, and a previously non-described formation of four Adriatic haplotypes (Balkan cluster). Haplotypes of the Balkan cluster and the other Adriatic haplotypes, do not represent a contiguous haplotype lineage and appear not to be closely related, indicating independent arrivals into the Adriatic drainage and suggesting successive colonization events. Despite the presence of marmoratus haplotypes in Albania, no marbled phenotype was found, confirming previously reported findings that there is no association between this phenotype and marmoratus haplotypes.
Molecular phylogenetics and evolution, 2010
Freshwater fauna of ancient lakes frequently contain endemic taxa thought to have originated duri... more Freshwater fauna of ancient lakes frequently contain endemic taxa thought to have originated during the long existence of these lakes, yet uncertainties remain as to whether they represent distinct genetic lineages with respect to more widespread relatives and to the relative roles of isolation and dispersal in their evolution. Phylogenetic analyses of sequence variation at nuclear and mitochondrial genes were used to examine these issues for the freshwater fish genus Barbus in two European ancient lake systems on the Balkan Peninsula. The nuclear and mitochondrial data yielded concordant phylogeographic patterns though incomplete sorting of nuclear haplotypes between some mitochondrial clades was detected. The distributions of two currently recognized species investigated here do not match the distributions of evolutionary lineages revealed by phylogenetic analyses. The Prespa barbel, Barbus prespensis, is not endemic to the lakes Prespa as previously thought but is instead found to be widespread in the southeastern Adriatic Sea basin, with a distribution largely corresponding to the basin of the now extinct Lake Maliq historically connected with Lake Prespa. On the other hand, a cryptic phylogenetic subdivision in a widespread species, B. rebeli, was discovered to be more distant from B. rebeli than from other Barbus species and to be endemic to the system of connected lakes Ohrid and Shkodra. The division coincides with the hydrogeographical boundary delimiting distributions of other freshwater fishes, and we suggest that this newly discovered evolutionary lineage represents a distinct species. These findings support the emerging pattern that endemic taxa have evolved not through isolation of individual lakes, but in systems of currently and historically interconnected lakes and their wider basins.
Molecular Phylogenetics and Evolution - MOL PHYLOGENET EVOL, 2010
Freshwater fauna of ancient lakes frequently contain endemic taxa thought to have originated duri... more Freshwater fauna of ancient lakes frequently contain endemic taxa thought to have originated during the long existence of these lakes, yet uncertainties remain as to whether they represent distinct genetic lineages with respect to more widespread relatives and to the relative roles of isolation and dispersal in their evolution. Phylogenetic analyses of sequence variation at nuclear and mitochondrial genes were used to examine these issues for the freshwater fish genus Barbus in two European ancient lake systems on the Balkan Peninsula. The nuclear and mitochondrial data yielded concordant phylogeographic patterns though incomplete sorting of nuclear haplotypes between some mitochondrial clades was detected. The distributions of two currently recognized species investigated here do not match the distributions of evolutionary lineages revealed by phylogenetic analyses. The Prespa barbel, Barbus prespensis, is not endemic to the lakes Prespa as previously thought but is instead found to be widespread in the southeastern Adriatic Sea basin, with a distribution largely corresponding to the basin of the now extinct Lake Maliq historically connected with Lake Prespa. On the other hand, a cryptic phylogenetic subdivision in a widespread species, B. rebeli, was discovered to be more distant from B. rebeli than from other Barbus species and to be endemic to the system of connected lakes Ohrid and Shkodra. The division coincides with the hydrogeographical boundary delimiting distributions of other freshwater fishes, and we suggest that this newly discovered evolutionary lineage represents a distinct species. These findings support the emerging pattern that endemic taxa have evolved not through isolation of individual lakes, but in systems of currently and historically interconnected lakes and their wider basins.
Genetics Selection Evolution, 2009
To determine the genetic architecture of trout in Albania, 87 individuals were collected from 19 ... more To determine the genetic architecture of trout in Albania, 87 individuals were collected from 19 riverine and lacustrine sites in Albania, FYROM and Greece. All individuals were analyzed for sequence variation in the mtDNA control region. Among fourteen haplotypes detected, four previously unpublished haplotypes, bearing a close relationship to haplotypes of the Adriatic and marmoratus lineages of Salmo trutta, were revealed. Ten previously described haplotypes, characteristic of S. ohridanus, S. letnica and the Adriatic and Mediterranean lineages of S. trutta, were also detected. Haplotypes detected in this study were placed in a well supported branch of S. ohridanus, and a cluster of Mediterranean -Adriatic -marmoratus haplotypes, which were further delimited into three subdivisions of Mediterranean, marmoratus, and a previously non-described formation of four Adriatic haplotypes (Balkan cluster). Haplotypes of the Balkan cluster and the other Adriatic haplotypes, do not represent a contiguous haplotype lineage and appear not to be closely related, indicating independent arrivals into the Adriatic drainage and suggesting successive colonization events. Despite the presence of marmoratus haplotypes in Albania, no marbled phenotype was found, confirming previously reported findings that there is no association between this phenotype and marmoratus haplotypes.
Melovski, D., Ivanov, Gj., Stojanov, A., Avukatov, V., Trajçe, A., Hoxha, B., von Arx, M., Breite... more Melovski, D., Ivanov, Gj., Stojanov, A., Avukatov, V., Trajçe, A., Hoxha, B., von Arx, M., Breitenmoser-Würsten, Ch., Hristovski, S., Shumka, S., Breitenmoser, U. (2013). Distribution and conservation status of the Balkan lynx (Lynx lynx balcanicus . Proceedings of the 4 th Congress of Ecologists of Macedonia with International Participation, Ohrid, 12-15 October 2012. Macedonian Ecological Society, Special issue 28, Skopje.