Aryeh Grossman | Midwestern University (original) (raw)

Papers by Aryeh Grossman

Research paper thumbnail of Functional Inference in Paleoanthropology: Theory and Practice

The Quarterly Review of Biology

Research paper thumbnail of Giant dwarf crocodiles from the Miocene of Kenya and crocodylid faunal dynamics in the late Cenozoic of East Africa

Research paper thumbnail of A Novel Description of Breast Anatomy Indicates the Need for New Surgical Approaches

Research paper thumbnail of Smith et al 2018 SVP Basilemys-01-01

Research paper thumbnail of A new baenid, "Trinitichelys" maini sp. nov., and other fossil turtles from the Upper Cretaceous Arlington Archosaur Site (Woodbine Formation, Cenomanian), Texas, USA

Palaeontologia Electronica, 2019

Research paper thumbnail of A Transitional Gundi (Rodentia: Ctenodactylidae) from the Miocene of Israel

PloS one, 2016

We describe a new species of gundi (Rodentia: Ctenodactylidae: Ctenodactylinae), Sayimys negevens... more We describe a new species of gundi (Rodentia: Ctenodactylidae: Ctenodactylinae), Sayimys negevensis, on the basis of cheek teeth from the Early Miocene of the Rotem Basin, southern Israel. The Rotem ctenodactylid differs from all known ctenodactylid species, including Sayimys intermedius, which was first described from the Middle Miocene of Saudi Arabia. Instead, it most resembles Sayimys baskini from the Early Miocene of Pakistan in characters of the m1-2 (e.g., the mesoflexid shorter than the metaflexid, the obliquely orientated hypolophid, and the presence of a strong posterolabial ledge) and the upper molars (e.g., the paraflexus that is longer than the metaflexus). However, morphological (e.g., presence of a well-developed paraflexus on unworn upper molars) and dimensional (regarding, in particular, the DP4 and M1 or M2) differences between the Rotem gundi and Sayimys baskini distinguish them and testify to the novelty and endemicity of the former. In its dental morphology, Say...

Research paper thumbnail of New Fauna from Loperot Contributes to the Understanding of Early Miocene Catarrhine Communities

International Journal of Primatology, 2014

The site of Loperot in West Turkana, Kenya, is usually assigned to the Early Miocene. Recent disc... more The site of Loperot in West Turkana, Kenya, is usually assigned to the Early Miocene. Recent discoveries at Loperot, including catarrhine primates, led to a revision of its mammalian fauna. Our revision of the fauna at Loperot shows an unusual taxonomic composition of the catarrhine community as well as several other unique mammalian taxa. Loperot shares two non-cercopithecoid catarrhine taxa with Early Miocene

Research paper thumbnail of Adaptations to cursoriality and digit reduction in the forelimb of the African wild dog (Lycaon pictus

PeerJ, 2020

Background: The African wild dog (Lycaon pictus), an endangered canid native to southern and east... more Background: The African wild dog (Lycaon pictus), an endangered canid native to southern and eastern Africa, is distinct among canids in being described as entirely tetradactyl and in its nomadic lifestyle and use of exhaustive predation to capture its prey instead of speed, strength, or stealth. These behavioral and morphological traits suggest a potentially unique set of adaptations. Methods: Here, we dissected the forelimbs of an adult male L. pictus specimen and performed detailed descriptions and quantitative analyses of the musculoskeletal anatomy. Results: Statistical comparisons of muscle masses and volumes revealed that L. pictus has relatively smaller wrist rotators (mm. pronator teres, pronator quadratus, supinator) than any other included carnivoran taxon, suggesting adaptive pressures for antebrachial stability over rotatory movement in the carpus of L. pictus. While a complete digit I is absent in L. pictus, a vestigial first metacarpal was discovered, resulting in changes to insertions of mm. extensor digiti I et II, abductor (et opponens) digiti I and flexor digiti I brevis. Mm. anconeus, brachialis and flexor carpi ulnaris caput ulnare all have more extensive origins in L. pictus than other canids suggesting an emphasis on posture and elbow stability. M. triceps brachii caput laterale has a larger origin in L. pictus and m. triceps brachii caput longum has an additional accessory head. Electromyographic studies have shown this muscle is active during the stance phase of trotting and galloping and is important for storing elastic energy during locomotion. We interpret these differences in size and attachments of muscles in L. pictus as adaptations for long distance running in this highly cursorial species, likely important for exhaustive predation. Absence of a full digit I in L. pictus may increase speed and stride length; however, the retention of a vestigial digit permits the attachment of reduced pollical muscles which may provide additional stability and proprioception to the carpus.

Research paper thumbnail of Reconstruction of the Early Miocene Critical Zone at Loperot, Southwestern Turkana, Kenya

Frontiers in Ecology and Evolution, 2019

The Hominoidea (apes and, eventually, humans) and Cercopithecoidea (Old World monkeys) diverged f... more The Hominoidea (apes and, eventually, humans) and Cercopithecoidea (Old World monkeys) diverged from a common ancestor during the late Oligocene (~25 Ma) in East Africa. Subsequently, both catarrhine groups diversified in the early Miocene (~23-16 Ma), making this time period an extremely important one for understanding our (deep) ancestral roots and the link between our hominoid ancestors and the environment. A remote region of west Turkana known as Loperot provides an exciting opportunity to study early Miocene primate paleoecology and has the potential to reveal the phylogenetic relationship between cercopithecoid monkeys and stem hominoids. The site of LpM4 is particularly fossiliferous and has produced a diverse fauna that includes several catarrhine fossils that await description. Radioisotopic dating indicates that the Loperot fauna are older than ~17 Ma (early Miocene). Using geochemical data from paleosols and stable isotope values from rhizolith calcite, coupled with detai...

Research paper thumbnail of Utilizing Stratigraphy and Stable Isotope Geochemistry to Reconstruct the Ancient Environments of Our Primate Ancestors: Loperot, Great Rift Valley, Kenya

Research paper thumbnail of Hindlimb anatomy of the African painted dog ( Lycaon pictus )

The African wild dog, Lycaon pictus, is classified as Endangered by the IUCN, with as few as 1400... more The African wild dog, Lycaon pictus, is classified as Endangered by the IUCN, with as few as 1400 mature individuals currently surviving in the wild, due primarily to habitat fragmentation. African...

Research paper thumbnail of Rhinocerotidae from the early Miocene of the Negev (Israel) and implications for the dispersal of early Neogene rhinoceroses

A revision of the rhinocerotid material from the Negev (Israel), dating back to the early Miocene... more A revision of the rhinocerotid material from the Negev (Israel), dating back to the early Miocene (MN3 in the European Mammal Biochronology), highlights the presence of Brachypotherium and a taxon close to Gaindatherium in the Levantine corridor. A juvenile mandible, investigated using CT scanning, displays morphologically distinct characters consistent with Brachypotherium cf. B. snowi rather than with other Eurasian representatives of this genus. Some postcranial remains from the Negev, such as a humerus, display features that distinguish it among Miocene taxa. We attribute these postcrania to cf. Gaindatherium sp., a taxon never recorded outside the Siwaliks until now. This taxon dispersed into the Levantine region during the late early Miocene, following a pattern similar to other South Asian taxa. Brachypotherium cf. B. snowi probably occurred in the Levantine region and then in North Africa during the early Miocene because its remains are known from slightly younger localities such as Moghara (Egypt) and Jebel Zelten (Libya). The occurrence cf. Gaindatherium sp. represents a previously unrecorded range expansion out of Southeast Asia. These new records demonstrate the paleogeographic importance of the Levantine region showcasing the complex role of the Levantine corridor in intercontinental dispersals between Asia and Europe as well as Eurasia and Africa.

Research paper thumbnail of An early bothremydid from the Arlington Archosaur Site of Texas

Scientific Reports

Four turtle taxa are previously documented from the Cenomanian Arlington Archosaur Site (AAS) of ... more Four turtle taxa are previously documented from the Cenomanian Arlington Archosaur Site (AAS) of the Lewisville Formation (Woodbine Group) in Texas. Herein, we describe a new side-necked turtle (Pleurodira), Pleurochayah appalachius gen. et sp. nov., which is a basal member of the Bothremydidae. Pleurochayah appalachius gen. et sp. nov. shares synapomorphic characters with other bothremydids, including shared traits with Kurmademydini and Cearachelyini, but has a unique combination of skull and shell traits. The new taxon is significant because it is the oldest crown pleurodiran turtle from North America and Laurasia, predating bothremynines Algorachelus peregrinus and Paiutemys tibert from Europe and North America respectively. This discovery also documents the oldest evidence of dispersal of crown Pleurodira from Gondwana to Laurasia. Pleurochayah appalachius gen. et sp. nov. is compared to previously described fossil pleurodires, placed in a modified phylogenetic analysis of pelo...

Research paper thumbnail of Rhinocerotidae from the early Miocene of the Negev (Israel) and implications for the dispersal of early Neogene rhinoceroses

Journal of Paleontology

A revision of the rhinocerotid material from the Negev (Israel), dating back to the early Miocene... more A revision of the rhinocerotid material from the Negev (Israel), dating back to the early Miocene (MN3 in the European Mammal Biochronology), highlights the presence of Brachypotherium and a taxon close to Gaindatherium in the Levantine corridor. A juvenile mandible, investigated using CT scanning, displays morphologically distinct characters consistent with Brachypotherium cf. B. snowi rather than with other Eurasian representatives of this genus. Some postcranial remains from the Negev, such as a humerus, display features that distinguish it among Miocene taxa. We attribute these postcrania to cf. Gaindatherium sp., a taxon never recorded outside the Siwaliks until now. This taxon dispersed into the Levantine region during the late early Miocene, following a pattern similar to other South Asian taxa. Brachypotherium cf. B. snowi probably occurred in the Levantine region and then in North Africa during the early Miocene because its remains are known from slightly younger localities...

Research paper thumbnail of Adaptations to cursoriality and digit reduction in the forelimb of the African wild dog (Lycaon pictus)

PeerJ

Background The African wild dog (Lycaon pictus), an endangered canid native to southern and easte... more Background The African wild dog (Lycaon pictus), an endangered canid native to southern and eastern Africa, is distinct among canids in being described as entirely tetradactyl and in its nomadic lifestyle and use of exhaustive predation to capture its prey instead of speed, strength, or stealth. These behavioral and morphological traits suggest a potentially unique set of adaptations. Methods Here, we dissected the forelimbs of an adult male L. pictus specimen and performed detailed descriptions and quantitative analyses of the musculoskeletal anatomy. Results Statistical comparisons of muscle masses and volumes revealed that L. pictus has relatively smaller wrist rotators (mm. pronator teres, pronator quadratus, supinator) than any other included carnivoran taxon, suggesting adaptive pressures for antebrachial stability over rotatory movement in the carpus of L. pictus. While a complete digit I is absent in L. pictus, a vestigial first metacarpal was discovered, resulting in change...

Research paper thumbnail of Adaptations to cursoriality and digit reduction in the forelimb of the African wild dog ( Lycaon pictus )

Research paper thumbnail of New Miocene Carnivora (Mammalia) from Moruorot and Kalodirr, Kenya

Palaeontologia electronica

Research paper thumbnail of Microanatomical Analyses (Paleohistology and Computed Tomography) Reveal Two Helochelydrid Taxa in the Woodbine Formation, Cenomanian, Texas

Research paper thumbnail of Shell Histology of the Stem Trionychians (Adocusia) Adocus Cf. Bossi and Nanhsiungchelyidae from the Cenomanian (Late Cretaceous) Arlington Archosaur Site, Woodbine Formation, Tarrant County, Texas, Usa

Research paper thumbnail of First record of Sivameryx (Cetartiodactyla, Anthracotheriidae) from the lower Miocene of Israel highlights the importance of the Levantine Corridor as a dispersal route between Eurasia and Africa

Journal of Vertebrate Paleontology

Research paper thumbnail of Functional Inference in Paleoanthropology: Theory and Practice

The Quarterly Review of Biology

Research paper thumbnail of Giant dwarf crocodiles from the Miocene of Kenya and crocodylid faunal dynamics in the late Cenozoic of East Africa

Research paper thumbnail of A Novel Description of Breast Anatomy Indicates the Need for New Surgical Approaches

Research paper thumbnail of Smith et al 2018 SVP Basilemys-01-01

Research paper thumbnail of A new baenid, "Trinitichelys" maini sp. nov., and other fossil turtles from the Upper Cretaceous Arlington Archosaur Site (Woodbine Formation, Cenomanian), Texas, USA

Palaeontologia Electronica, 2019

Research paper thumbnail of A Transitional Gundi (Rodentia: Ctenodactylidae) from the Miocene of Israel

PloS one, 2016

We describe a new species of gundi (Rodentia: Ctenodactylidae: Ctenodactylinae), Sayimys negevens... more We describe a new species of gundi (Rodentia: Ctenodactylidae: Ctenodactylinae), Sayimys negevensis, on the basis of cheek teeth from the Early Miocene of the Rotem Basin, southern Israel. The Rotem ctenodactylid differs from all known ctenodactylid species, including Sayimys intermedius, which was first described from the Middle Miocene of Saudi Arabia. Instead, it most resembles Sayimys baskini from the Early Miocene of Pakistan in characters of the m1-2 (e.g., the mesoflexid shorter than the metaflexid, the obliquely orientated hypolophid, and the presence of a strong posterolabial ledge) and the upper molars (e.g., the paraflexus that is longer than the metaflexus). However, morphological (e.g., presence of a well-developed paraflexus on unworn upper molars) and dimensional (regarding, in particular, the DP4 and M1 or M2) differences between the Rotem gundi and Sayimys baskini distinguish them and testify to the novelty and endemicity of the former. In its dental morphology, Say...

Research paper thumbnail of New Fauna from Loperot Contributes to the Understanding of Early Miocene Catarrhine Communities

International Journal of Primatology, 2014

The site of Loperot in West Turkana, Kenya, is usually assigned to the Early Miocene. Recent disc... more The site of Loperot in West Turkana, Kenya, is usually assigned to the Early Miocene. Recent discoveries at Loperot, including catarrhine primates, led to a revision of its mammalian fauna. Our revision of the fauna at Loperot shows an unusual taxonomic composition of the catarrhine community as well as several other unique mammalian taxa. Loperot shares two non-cercopithecoid catarrhine taxa with Early Miocene

Research paper thumbnail of Adaptations to cursoriality and digit reduction in the forelimb of the African wild dog (Lycaon pictus

PeerJ, 2020

Background: The African wild dog (Lycaon pictus), an endangered canid native to southern and east... more Background: The African wild dog (Lycaon pictus), an endangered canid native to southern and eastern Africa, is distinct among canids in being described as entirely tetradactyl and in its nomadic lifestyle and use of exhaustive predation to capture its prey instead of speed, strength, or stealth. These behavioral and morphological traits suggest a potentially unique set of adaptations. Methods: Here, we dissected the forelimbs of an adult male L. pictus specimen and performed detailed descriptions and quantitative analyses of the musculoskeletal anatomy. Results: Statistical comparisons of muscle masses and volumes revealed that L. pictus has relatively smaller wrist rotators (mm. pronator teres, pronator quadratus, supinator) than any other included carnivoran taxon, suggesting adaptive pressures for antebrachial stability over rotatory movement in the carpus of L. pictus. While a complete digit I is absent in L. pictus, a vestigial first metacarpal was discovered, resulting in changes to insertions of mm. extensor digiti I et II, abductor (et opponens) digiti I and flexor digiti I brevis. Mm. anconeus, brachialis and flexor carpi ulnaris caput ulnare all have more extensive origins in L. pictus than other canids suggesting an emphasis on posture and elbow stability. M. triceps brachii caput laterale has a larger origin in L. pictus and m. triceps brachii caput longum has an additional accessory head. Electromyographic studies have shown this muscle is active during the stance phase of trotting and galloping and is important for storing elastic energy during locomotion. We interpret these differences in size and attachments of muscles in L. pictus as adaptations for long distance running in this highly cursorial species, likely important for exhaustive predation. Absence of a full digit I in L. pictus may increase speed and stride length; however, the retention of a vestigial digit permits the attachment of reduced pollical muscles which may provide additional stability and proprioception to the carpus.

Research paper thumbnail of Reconstruction of the Early Miocene Critical Zone at Loperot, Southwestern Turkana, Kenya

Frontiers in Ecology and Evolution, 2019

The Hominoidea (apes and, eventually, humans) and Cercopithecoidea (Old World monkeys) diverged f... more The Hominoidea (apes and, eventually, humans) and Cercopithecoidea (Old World monkeys) diverged from a common ancestor during the late Oligocene (~25 Ma) in East Africa. Subsequently, both catarrhine groups diversified in the early Miocene (~23-16 Ma), making this time period an extremely important one for understanding our (deep) ancestral roots and the link between our hominoid ancestors and the environment. A remote region of west Turkana known as Loperot provides an exciting opportunity to study early Miocene primate paleoecology and has the potential to reveal the phylogenetic relationship between cercopithecoid monkeys and stem hominoids. The site of LpM4 is particularly fossiliferous and has produced a diverse fauna that includes several catarrhine fossils that await description. Radioisotopic dating indicates that the Loperot fauna are older than ~17 Ma (early Miocene). Using geochemical data from paleosols and stable isotope values from rhizolith calcite, coupled with detai...

Research paper thumbnail of Utilizing Stratigraphy and Stable Isotope Geochemistry to Reconstruct the Ancient Environments of Our Primate Ancestors: Loperot, Great Rift Valley, Kenya

Research paper thumbnail of Hindlimb anatomy of the African painted dog ( Lycaon pictus )

The African wild dog, Lycaon pictus, is classified as Endangered by the IUCN, with as few as 1400... more The African wild dog, Lycaon pictus, is classified as Endangered by the IUCN, with as few as 1400 mature individuals currently surviving in the wild, due primarily to habitat fragmentation. African...

Research paper thumbnail of Rhinocerotidae from the early Miocene of the Negev (Israel) and implications for the dispersal of early Neogene rhinoceroses

A revision of the rhinocerotid material from the Negev (Israel), dating back to the early Miocene... more A revision of the rhinocerotid material from the Negev (Israel), dating back to the early Miocene (MN3 in the European Mammal Biochronology), highlights the presence of Brachypotherium and a taxon close to Gaindatherium in the Levantine corridor. A juvenile mandible, investigated using CT scanning, displays morphologically distinct characters consistent with Brachypotherium cf. B. snowi rather than with other Eurasian representatives of this genus. Some postcranial remains from the Negev, such as a humerus, display features that distinguish it among Miocene taxa. We attribute these postcrania to cf. Gaindatherium sp., a taxon never recorded outside the Siwaliks until now. This taxon dispersed into the Levantine region during the late early Miocene, following a pattern similar to other South Asian taxa. Brachypotherium cf. B. snowi probably occurred in the Levantine region and then in North Africa during the early Miocene because its remains are known from slightly younger localities such as Moghara (Egypt) and Jebel Zelten (Libya). The occurrence cf. Gaindatherium sp. represents a previously unrecorded range expansion out of Southeast Asia. These new records demonstrate the paleogeographic importance of the Levantine region showcasing the complex role of the Levantine corridor in intercontinental dispersals between Asia and Europe as well as Eurasia and Africa.

Research paper thumbnail of An early bothremydid from the Arlington Archosaur Site of Texas

Scientific Reports

Four turtle taxa are previously documented from the Cenomanian Arlington Archosaur Site (AAS) of ... more Four turtle taxa are previously documented from the Cenomanian Arlington Archosaur Site (AAS) of the Lewisville Formation (Woodbine Group) in Texas. Herein, we describe a new side-necked turtle (Pleurodira), Pleurochayah appalachius gen. et sp. nov., which is a basal member of the Bothremydidae. Pleurochayah appalachius gen. et sp. nov. shares synapomorphic characters with other bothremydids, including shared traits with Kurmademydini and Cearachelyini, but has a unique combination of skull and shell traits. The new taxon is significant because it is the oldest crown pleurodiran turtle from North America and Laurasia, predating bothremynines Algorachelus peregrinus and Paiutemys tibert from Europe and North America respectively. This discovery also documents the oldest evidence of dispersal of crown Pleurodira from Gondwana to Laurasia. Pleurochayah appalachius gen. et sp. nov. is compared to previously described fossil pleurodires, placed in a modified phylogenetic analysis of pelo...

Research paper thumbnail of Rhinocerotidae from the early Miocene of the Negev (Israel) and implications for the dispersal of early Neogene rhinoceroses

Journal of Paleontology

A revision of the rhinocerotid material from the Negev (Israel), dating back to the early Miocene... more A revision of the rhinocerotid material from the Negev (Israel), dating back to the early Miocene (MN3 in the European Mammal Biochronology), highlights the presence of Brachypotherium and a taxon close to Gaindatherium in the Levantine corridor. A juvenile mandible, investigated using CT scanning, displays morphologically distinct characters consistent with Brachypotherium cf. B. snowi rather than with other Eurasian representatives of this genus. Some postcranial remains from the Negev, such as a humerus, display features that distinguish it among Miocene taxa. We attribute these postcrania to cf. Gaindatherium sp., a taxon never recorded outside the Siwaliks until now. This taxon dispersed into the Levantine region during the late early Miocene, following a pattern similar to other South Asian taxa. Brachypotherium cf. B. snowi probably occurred in the Levantine region and then in North Africa during the early Miocene because its remains are known from slightly younger localities...

Research paper thumbnail of Adaptations to cursoriality and digit reduction in the forelimb of the African wild dog (Lycaon pictus)

PeerJ

Background The African wild dog (Lycaon pictus), an endangered canid native to southern and easte... more Background The African wild dog (Lycaon pictus), an endangered canid native to southern and eastern Africa, is distinct among canids in being described as entirely tetradactyl and in its nomadic lifestyle and use of exhaustive predation to capture its prey instead of speed, strength, or stealth. These behavioral and morphological traits suggest a potentially unique set of adaptations. Methods Here, we dissected the forelimbs of an adult male L. pictus specimen and performed detailed descriptions and quantitative analyses of the musculoskeletal anatomy. Results Statistical comparisons of muscle masses and volumes revealed that L. pictus has relatively smaller wrist rotators (mm. pronator teres, pronator quadratus, supinator) than any other included carnivoran taxon, suggesting adaptive pressures for antebrachial stability over rotatory movement in the carpus of L. pictus. While a complete digit I is absent in L. pictus, a vestigial first metacarpal was discovered, resulting in change...

Research paper thumbnail of Adaptations to cursoriality and digit reduction in the forelimb of the African wild dog ( Lycaon pictus )

Research paper thumbnail of New Miocene Carnivora (Mammalia) from Moruorot and Kalodirr, Kenya

Palaeontologia electronica

Research paper thumbnail of Microanatomical Analyses (Paleohistology and Computed Tomography) Reveal Two Helochelydrid Taxa in the Woodbine Formation, Cenomanian, Texas

Research paper thumbnail of Shell Histology of the Stem Trionychians (Adocusia) Adocus Cf. Bossi and Nanhsiungchelyidae from the Cenomanian (Late Cretaceous) Arlington Archosaur Site, Woodbine Formation, Tarrant County, Texas, Usa

Research paper thumbnail of First record of Sivameryx (Cetartiodactyla, Anthracotheriidae) from the lower Miocene of Israel highlights the importance of the Levantine Corridor as a dispersal route between Eurasia and Africa

Journal of Vertebrate Paleontology