Coleoptera (original) (raw)
For the scientific journal named Coleoptera, see ISSN 0945-1889.
Superregnum: Eukaryota
Cladus: Podiata
Cladus: Amorphea
Cladus: Obazoa
Cladus: Opisthokonta
Cladus: Holozoa
Cladus: Filozoa
Cladus: Choanozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: ParaHoxozoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Superphylum: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Cladus: Mandibulata
Cladus: Pancrustacea
Cladus: Allotriocarida
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Holometabola
Superordo: Coleopterida
Ordo: Coleoptera
Subordines (4 + 1†):Adephaga –Archostemata –Myxophaga –Polyphaga – †Alphacoleoptera
Coleoptera incertae sedis:
Familiae: †Ponomarenkiumidae
Genera: †_Anepismus_ – †_Anhuistoma_ – †_Apheloodes_ – †_Apistotes_ – †_Archosyne_ – †_Babuskaya_ – †_Biadelater_ – †_Byrrhydium_ – †_Cardiosyne_ – †_Cistelites_ – †_Cretaceites_ – †_Dysarestus_ – †_Elaterina_ – †_Enamma_ – †_Fengningia_ – †_Fuscicupes_ – †_Gemelina_ – †_Glaphyropterites_ – †_Grahamelytron_ – †_Helophoropsis_ – †_Hydrophilites_ – †_Keleusticus_ – †_Leioodes_ – †_Leptopodocoleus_ – †_Megacentrus_ – †_Micrelaterium_ – †_Microentomus_ – †_Mimelater_ – †_Parabuprestium_ – †_Paragrilium_ – †_Peltocoleops_ – †_Plastelater_ – †_Praedodromeus_ – †_Pseudoelateropsis_ – †_Stenelytron_ – †_Toksunius_ – †_Trialarva_ – †_Tychon_ – †_Xinbinia_ – †_Yanqingia_ – †_Zetemenos_
Species: †_"Brochocoleus" indibili_
Collective groups: †_Coleopteron_
Overview of familiae (186 + †45)
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Acanthocnemidae –Aderidae –Agapythidae –Agyrtidae –Akalyptoischiidae –Alexiidae –Amphizoidae –Anamorphidae –Anthicidae –Anthribidae –Archeocrypticidae –Artematopodidae –Aspidytidae –Attelabidae –Badmaateridae –Belidae –Biphyllidae –Boganiidae –Boridae –Bostrichidae –Bothrideridae –Brachyceridae –Brachypsectridae –Brentidae –Buprestidae –Byrrhidae –Byturidae –Callirhipidae –Cantharidae –Carabidae –Caridae –Cavognathidae –Cerambycidae –Cerasommatidiidae –Cerophytidae –Cerylonidae –Chaetosomatidae –Chalcodryidae –Chelonariidae –Chrysomelidae –Cicindelidae –Ciidae –Clambidae –Cleridae –Cneoglossidae –Coccinellidae –Corylophidae –Crowsoniellidae –Cryptophagidae –Cucujidae –Cupedidae –Curculionidae –Cybocephalidae –Cyclaxyridae –Dascillidae –Decliniidae –Dermestidae –Derodontidae –Discolomatidae –Disteniidae –Dryophthoridae –Dryopidae –Dytiscidae –Elateridae –Elmidae –Endecatomidae –Endomychidae –Epimetopidae –Erotylidae –Eucinetidae –Eucnemidae –Eulichadidae –Eupsilobiidae –Euxestidae –Georissidae –Gyrinidae –Haliplidae –Helophoridae –Helotidae –Heteroceridae –Histeridae –Hobartiidae –Hydraenidae –Hydrochidae –Hydrophilidae –Hydroscaphidae –Hygrobiidae –Hylecoetidae –Ischaliidae –Jacobsoniidae –Jurodidae –Kateretidae –Laemophloeidae –Lagrioididae –Lamingtoniidae –Lampyridae –Latridiidae –Leiodidae –Lepiceridae –Lophocateridae –Limnichidae –Lutrochidae –Lycidae –Lymexylidae –Mauroniscidae –Megalopodidae –Melandryidae –Meloidae –Melyridae –Meruidae –Micromalthidae –Monotomidae –Mordellidae –Murmidiidae –Mycetaeidae –Mycetophagidae –Mycteridae –Myraboliidae –Nemonychidae –Nitidulidae –Nosodendridae –Noteridae –Oedemeridae –Omalisidae –Omethidae –Ommatidae –Orsodacnidae –Oxypeltidae –Passandridae –Peltidae –Phalacridae –Phengodidae –Phloeostichidae –Phloiophilidae –Phycosecidae –Plastoceridae –Priasilphidae –Prionoceridae –Promecheilidae –Prostomidae –Protelmidae –Protocucujidae –Protopeltidae –Psephenidae –Pterogeniidae –Ptiliidae –Ptilodactylidae –Ptinidae –Pyrochroidae –Pythidae –Rhadalidae –Rentoniidae –Rhagophthalmidae –Rhinorhipidae –Rhipiceridae –Ripiphoridae –Salpingidae –Schizopodidae –Scirtidae –Scraptiidae –Silphidae –Silvanidae –Smicripidae –Spercheidae –Sphaeritidae –Sphaeriusidae –Sphindidae –Staphylinidae –Stenotrachelidae –Synchroidae –Synteliidae –Tasmosalpingidae –Telegeusidae –Tenebrionidae –Teredidae –Tetratomidae –Thanerocleridae –Throscidae –Thymalidae –Torridincolidae –Trachypachidae –Trictenotomidae –Trogossitidae –Ulodidae –Vesperidae –Zopheridae
Fossil: †Ademosynidae – †Apophisandridae – †Asiocoleidae – †Bellimordellidae – †Berendtimiridae – †Boleopsidae – †Catiniidae – †Coleopsidae – †Colymbotethidae – †Coptoclavidae – †Elodophthalmidae – †Labradorocoleidae – †Lasiosynidae – †Liadytidae – †Liaoximordellidae – †Lithoscarabaeidae – †Magnocoleidae – †Mesocinetidae – †Mirimordellidae – †Oborocoleidae – †Obrieniidae – †Parahygrobiidae – †Paralucanidae – †Parandrexidae – †Peltosynidae – †Permocupedidae – †Permosynidae – †Phoroschizidae – †Ponomarenkiumidae – †Praelateriidae – †Praemordellidae – †Rhombocoleidae – †Schizocoleidae – †Taldycupedidae – †Tetrameropseidae – †Tshekardocoleidae – †Triadocupedidae – †Triamyxidae – †Triaplidae – †Tritarsusidae – †Ulyanidae – †Wabbelidae – †Wuhuidae
Ichnofamiliae: †Coprinisphaeridae – †Pallichnidae
Coleoptera Linnaeus, 1758: 341
- Eleuterata Fabricius, 1775: 1
- Fabricius, J.C. 1775. Systema entomologiae, sistens insectorum classes, ordines, genera, species, adiectis synonymis, locis, descriptionibus, observationibus. Libraria Kortii, Flensburgi et Lipsiae. xvi+ 832 pp. DOI: 10.5962/bhl.title.36510. BHL. Reference page.
- Linnaeus, C. 1758. Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio Decima, Reformata. Tomus I. Holmiæ (Stockholm): impensis direct. Laurentii Salvii. 824 pp. DOI: 10.5962/bhl.title.542 BHL Reference page.
Additional references
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Nomenclature and classification
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- Bouchard, P., Bousquet, Y., Davies, A.E., Alonso-Zarazaga, M.A., Lawrence, J.F., Lyal, C.H.C., Newton, A.F., Reid, C.A.M., Schmitt, M., Ślipiński, S.A. & Smith, A.B.T. 2011. Family-group names in Coleoptera (Insecta). ZooKeys 88: 1–972. DOI: 10.3897/zookeys.88.807
. PMC: 3088472
. Reference page. - Bouchard, P., Bousquet, Y., Davies, A.E. & Cai, C-Y. 2024. On the nomenclatural status of type genera in Coleoptera (Insecta). ZooKeys 1194: 1–984. DOI: 10.3897/zookeys.1194.106440
Reference page. - Crotch, G.R. 1871. List of all the Coleoptera described A.D. 1758-1821, referred to their modern genera. Cambridge. [1] + 24 pp.
- Lawrence, J.F., Beutel, R.G., Leschen, R.A.B. & Ślipiński, A. 2010. 1. Changes in classification and list of families and subfamilies. Pp. 1–7. In: Leschen, R.A.B., Beutel, R.G. & Lawrence, J.F. (volume eds.) Coleoptera, beetles. Volume 2: Morphology and Systematics (Elateroidea, Bostrichiformia, Cucujiformia partim). In: Kristensen, N.P. & Beutel, R.G. (eds.) Handbook of Zoology. A natural history of the phyla of the animal kingdom. Volume IV. Arthropoda: Insecta. Part 38. Berlin, New York: Walter de Gruyter. ISBN 3110190753 ISBN 9783110190755. DOI: 10.1515/9783110911213.1
. - Lawrence, J.F. & Newton, A.F., Jr. 1982. Evolution and classification of beetles. Annual Review of Ecology and Systematics 13: 261–290. DOI: 10.1146/annurev.es.13.110182.001401
. Reference page. - Lawrence, J.F. & Newton, A.F. 1995. Families and subfamilies of Coleoptera (with selected genera, notes, references and data on family-group names). Pp. 779–1006. In: Pakaluk, J. & Ślipiński, S.A. (eds.) Biology, Phylogeny, and Classification of Coleoptera: Papers Celebrating the 80th Birthday of Roy A. Crowson. Volume 2. Museum i Instytut Zoologii PAN, Warszawa. x + 1092 pp. in 2 vols. ISBN 83-85192-34-4. ResearchGate
. Reference page. - Lawrence, J.F., Ślipiński, S.A. & Pakaluk, J. 1995. From Latreille to Crowson: a history of the higher-level classification of beetles. Pp. 87–154. in Pakaluk, J. & Ślipiński, S.A. (eds.) Biology, phylogeny, and classification of Coleoptera: papers celebrating the 80th birthday of Roy A. Crowson. Museum i Instytut Zoologii PAN, Warszawa.
- Slipinski, S.A., Leschen, R.A.B. & Lawrence, J.F. 2011. Order Coleoptera Linnaeus, 1758. Pp 203–208 In Zhang, Z.-Q. (ed.) 2011. Animal biodiversity: an outline of higher-level classification and survey of taxonomic richness. Zootaxa 3148: 1–237. Open access. Reference page. . (PDF) Reference page.
- Watt, J.C. 1975. Notes on priority of family-group names in Coleoptera. The Coleopterists Bulletin 29(1): 31–34. BHL.
- Beutel, R.G., Ge, S.-Q. & Hörnschemeyer, T. 2007 (Online) 2008 (Print). On the head morphology of Tetraphalerus, the phylogeny of Archostemata and the basal branching events in Coleoptera. Cladistics 24(3): 270–298. DOI: 10.1111/j.1096-0031.2007.00186.x
. Reference page. - Beutel, R.G. & Haas, F. 2000. Phylogenetic relationships of the suborders of Coleoptera (Insecta). Cladistics 16(1): 103–141. DOI: 10.1111/j.1096-0031.2000.tb00350.x
. Reference page. - Beutel, R.G. & Haas, F. 2003. Basal splitting events in Coleoptera. Pp. 160–161. In: Klass, K.D. (ed.) Proceedings of the 1st Dresden meeting on insect phylogeny: “Phylogenetic relationships within the insect orders” (Dresden, September 19–21, 2003). Entomologische Abhandlungen 61: 119–172. PDF. Reference page.
- Bocak, L., Barton, C., Crampton-Blatt, A., Chesters, D., Ahrens, D. & Vogler, A.P. 2014. Building the Coleoptera tree-of-life for >8000 species: composition of public DNA data and fit with Linnaean classification. Systematic Entomology 39(1): 97–110. DOI: 10.1111/syen.12037
. Reference page. - Boudinot, B.E., Fikáček, M., Lieberman, Z.E., Kusy, D., Bocak, L., Mckenna, D.D. & Beutel, R.G. 2023 (Online 2022). Systematic bias and the phylogeny of Coleoptera—A response to Cai et al. (2022) following the responses to Cai et al. (2020). Systematic Entomology 48(2): 223–232. DOI: 10.1111/syen.12570
. ResearchGate
. Reference page. - Boudinot, B.E., Yan, E.V., Prokop, J., Luo, X.-Z. & Beutel, R.G. 2023 (Online 2022). Permian parallelisms: reanalysis of †Tshekardocoleidae sheds light on the earliest evolution of the Coleoptera. Systematic Entomology 48(1): 69–96. DOI: 10.1111/syen.12562
. Reference page. - Cai, C., Tihelka, E., Giacomelli, M., Lawrence, J.F., Ślipiński, A., Kundrata, R., Yamamoto, S., Thayer, M.K., Newton, A.F., Leschen, R.A.B., Gimmel, M.L., Lü, L., Engel, M.S., Bouchard, P., Huang, D., Pisani, D. & Donoghue, P.C.J. 2022. Integrated phylogenomics and fossil data illuminate the evolution of beetles. Royal Society Open Science 9: 211771. DOI: 10.1098/rsos.211771
. [Supplemental material] Reference page. - Caterino, M.S., Shull, V.L., Hammond, P.M. & Vogler, A.P. 2002. Basal relationships of Coleoptera inferred from 18S rDNA sequences. Zoologica Scripta 31(1): 41–49. DOI: 10.1046/j.0300-3256.2001.00092.x
. Reference page. - Friedrich, F. & Beutel, R.G. 2006. The pterothoracic skeletomuscular system of Scirtoidea (Coleoptera: Polyphaga) and its implications for the high-level phylogeny of beetles. Journal of Zoological Systematics and Evolutionary Research 44(4): 290–315. DOI: 10.1111/j.1439-0469.2006.00369.x
. Reference page. - Friedrich, F., Farrell, B.D. & Beutel, R.G. 2008 (Online) 2009 (Print). The thoracic morphology of Archostemata and the relationships of the extant suborders of Coleoptera (Hexapoda). Cladistics 25(1): 1–37. DOI: 10.1111/j.1096-0031.2008.00233.x
. [Erratum: 2009. 25(2): 209. DOI: 10.1111/j.1096-0031.2009.00257.x
.] Reference page. - Hunt, T., Bergsten, J., Levkanicova, Z., Papadopoulou, A., St. John, O., Wild, R., Hammond, P.M., Ahrens, D., Balke, M., Caterino, M.S., Gómez-Zurita, J., Ribera, I., Barraclough, T.G., Bocáková, M., Bocak, L. & Vogler, A.P. 2007. A comprehensive phylogeny of beetles reveals the evolutionary origins of a superradiation. Science 318: 1913–1916. DOI: 10.1126/science.1146954
. Reference page. - Lawrence, J.F., Ślipiński, A., Seago, A.E., Thayer, M.K., Newton, A.F. & Marvaldi, A.E. 2011. Phylogeny of the Coleoptera based on morphological characters of adults and larvae. Annales Zoologici 61(1): 1–217. DOI: 10.3161/000345411X576725
. Reference page. - Vogler, A.P. & Caterino, M.S. 2003. The basal relationships of Coleoptera based on 18S rRNA sequences. Pp. 161–162. In: Klass, K.D. (ed.) Proceedings of the 1st Dresden meeting on insect phylogeny: “Phylogenetic relationships within the insect orders” (Dresden, September 19–21, 2003). Entomologische Abhandlungen 61: 119–172. PDF. Reference page.
Checklists and catalogues
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Abdel-Dayem, M.S., Abu El-Ghiet, U.M., El-Sheikh, T.M., Elgharbawy, A.A., Al-Fifi, Z.I.A. & Aldhafer, H.M. 2020. The first survey of the beetles (Coleoptera) of the Farasan Archipelago of the southern Red Sea, Kingdom of Saudi Arabia. ZooKeys, 959: 17–86. DOI: 10.3897/zookeys.959.51224
Reference page.Bousquet, Y., Bouchard, P., Davies, A.E. & Sikes, D.S. 2013. Checklist of beetles (Coleoptera) of Canada and Alaska. Second edition. ZooKeys 360: 1–44. DOI: 10.3897/zookeys.360.4742
. Reference page.Egorov, L.V., Ruchin, A.B., Semenov, V.B., Semionenkov, O.I. & Semishin, G.B. 2020. Checklist of the Coleoptera of Mordovia State Nature Reserve, Russia. ZooKeys 962: 13–122. DOI: 10.3897/zookeys.962.54477
Reference page.Gerlach, J. (ed.) 2009. The Coleoptera of the Seychelles islands. Series Faunistica 88. Sofia-Moscow: Pensoft Publishers. 266 pp. Reference page.
Hájek, J. & Bezděk, J. 2019. Annotated catalogue of beetles (Coleoptera) of the Socotra Archipelago. Zootaxa 4715(1): 1–76. DOI: 10.11646/zootaxa.4715.1.1
Reference page.Lamelas-López, L., Raposeiro, P.M., Borges, P.A.V. & Florencio, M. 2017. Annotated checklist of aquatic beetles (Coleoptera) and true bugs (Heteroptera) in the Azores Islands: new records and corrections of colonization status. Zootaxa 4353(1): 117–132. DOI: 10.11646/zootaxa.4353.1.7. Reference page.
Maddison, P. (compiler) 2010. Order Coleoptera. Pp. 409–436. In: Checklist of New Zealand Hexapoda. Pp. 396–467. In:
Macfarlane, R.P., Maddison, P.A., Andrew, I.G., Berry, J.A., Johns, P.M., Hoare, R.J.B., Larivière, M.-C., Greenslade, P., Henderson, R.C., Smithers, C.N., Palma, R.L., Ward, J.B., Pilgrim, R.L.C., Towns, D.R., McLellan, I., Teulon, D.A.J., Hitchings, T.R., Eastop, V.F., Martin, N.A., Fletcher, M.J., Stufkens, M.A.W., Dale, P.J., Burckhardt, D., Buckley, T.R. & Trewick, S.A. 2010. Phylum Arthropoda subphylum Hexapoda: Protura, springtails, Diplura, and insects. Pp. 233–467 in: Gordon, D.P. (ed.) 2010. New Zealand Inventory of Biodiversity. Volume 2. Kingdom Animalia. Chaetognatha, Ecdysozoa, ichnofossils. Canterbury University Press, Christchurch, New Zealand. Reference page. PDF. Reference page.
Majka, C.G. & Sikes, D.S. 2009. Thomas L. Casey and Rhode Island's precinctive beetles: taxonomic lessons and the utility of distributional checklists. In: Majka, C.G. & Klimaszewski, J. (eds.) Biodiversity, biosystematics, and ecology of Canadian Coleoptera II. ZooKeys 22: 267–283. DOI: 10.3897/zookeys.22.93
. Reference page.Tamutis, V., Tamutė, B. & Ferenca, R. 2011. A catalogue of Lithuanian beetles (Insecta, Coleoptera). ZooKeys 121: 1–494. DOI: 10.3897/zookeys.121.732
. Reference page.Traylor, C.R., Ulyshen, M.D., Cornish, J.W., Tigreros, G. & McHugh, J.V. 2025. Progress toward a list of saproxylic beetles (Coleoptera) in the southeastern USA. ZooKeys 1232: 1–95. DOI: 10.3897/zookeys.1232.143989
Reference page.Webster, R.P., Hughes, C. & Sweeney, J.D. 2022. The Coleoptera of the Province of Prince Edward Island, Canada: 295 new records from Lindgren funnel traps and a checklist to species. Zookeys 1107ː 1–158. DOI: 10.3897/zookeys.1107.82976
Reference page.Zaragoza-Caballero, S., Rodríguez-Mirón, G.M., Vega-Badillo, V., González-Ramírez, M., Zurita-García, M.L., Domínguez-León, D.E., López-Pérez, S., Gutiérrez-Carranza, I.G., Cifuentes-Ruiz, P., Pérez-Hernández, C.X., Ramírez del Valle, E. & Gutiérrez, N. 2019. A checklist of the Coleoptera (Insecta) from Morelos, Mexico. Zootaxa 4580(1): 1–122. DOI: 10.11646/zootaxa.4580.1.1
Reference page.Alekseev, V.I. 2013. The beetles (Insecta: Coleoptera) of Baltic amber: the checklist of described species and preliminary analysis of biodiversity. Zoology and Ecology 23(1): 5–12. DOI: 10.1080/21658005.2013.769717
. Reference page.Alekseev, V.I. 2017. A new coleopterous family Wabbelidae fam. nov. (Coleoptera: Cucujoidea) from Baltic amber (Cenozoic, Paleogene, Eocene). Baltic Journal of Coleopterology 17(1): 29–41. PDF. Reference page.
Batelka, J., Engel, M.S. & Prokop, J. 2018. A remarkable diversity of parasitoid beetles (Ripiphoridae) in Cretaceous amber, with a summary of the Mesozoic record of Tenebrionoidea. Cretaceous Research 90: 296–310. DOI: 10.1016/j.cretres.2018.04.019
. Reference page.Béthoux, O. 2009. The earliest beetle identified. Journal of Paleontology 83(6): 931–937. DOI: 10.1666/08-158.1
.Fikáček, M., Prokin, A., Angus, R.B., Ponomarenko, A., Yue, Y., Ren, D. & Prokop, J. 2012. Revision of Mesozoic fossils of the helophorid lineage of the superfamily Hydrophiloidea (Coleoptera: Polyphaga). Acta Entomologica Musei Nationalis Pragae 52(1): 89–127. PDF Reference page.
Jin, Z., Ślipiński, A., Pang, H. & Ren, D. 2013. A new Mesozoic species of soft-bodied plant beetle (Coleoptera: Dascillidae) from the Early Cretaceous of Inner Mongolia, China with a review of fossil Dascillidae. Annales Zoologici 63(3): 501–509. DOI: 10.3161/000345413X672492
. Reference page.Kirejtshuk, A.G. 2020. Taxonomic Review of Fossil Coleopterous Families (Insecta, Coleoptera). Suborder Archostemata: Superfamilies Coleopseoidea and Cupedoidea. Geosciences 10(2): 73. DOI: 10.3390/geosciences10020073
. Reference page.Kirejtshuk, A.G. & Ansorge, J. 2022 (Online) 2023 (Print). An extraordinarily preserved new genus and species of Trachypachidae (Coleoptera, Adephaga) from the Early Jurassic of Germany and a review of fossil trachypachid genera. Historical Biology 35(6): 958–975. DOI: 10.1080/08912963.2022.2071709
. Reference page.Kirejtshuk, A.G. & Prokin, A.A. 2018. The Position of the Palaeozoic Genus Tunguskagyrus Yan, Beutel et Lawrence in the Family Triaplidae sensu n. (Coleoptera, Archostemata: Schizophoroidea). Entomological Review 98: 872–882. DOI: 10.1134/S0013873818070084
. Reference page.Kundrata, R., Packova, G. & Hoffmannova, J. 2020. Fossil Genera in Elateridae (Insecta, Coleoptera): A Triassic Origin and Jurassic Diversification. Insects 11(6): 394. DOI: 10.3390/insects11060394
. Reference page.Ponomarenko, A.G. 1995. The geological history of beetles. Pp. 155–171. in Pakaluk, J. & Ślipiński, S.A. (eds.) Biology, phylogeny, and classification of Coleoptera: papers celebrating the 80th birthday of Roy A. Crowson. Museum i Instytut Zoologii PAN, Warszawa.
Ponomarenko, A.G. 2002. 2.2.1.3.2. Superorder Scarabaeidea Laicharting, 1781. Order Coleoptera Linné, 1758. The beetles. Pp. 164–176. In: Rasnitsyn, A.P. & Quicke, D.L.J. (eds.) History of Insects. Kluwer Academic Publishers, New York, Boston, Dordrecht, London, Moscow. ISBN 1-4020-0026-X (print), ISBN 0-306-47577-4 (e-book). DOI: 10.1007/0-306-47577-4
. Reference page.Ponomarenko, A.G. 2003. Ecological evolution of beetles (Insecta: Coleoptera). Acta zoologica cracoviensia 46(suppl.-Fossil Insects): 319–328. [1].
Ponomarenko, A.G. 2011. [New beetles (Insecta, Coleoptera) from Vyazniki Locality, Terminal Permian of European Russia]. Paleontologicheskii Zhurnal 2011(4): 55–63. [in Russian, English translation in Paleontological Journal 45(4): 414–422. DOI: 10.1134/S0031030111040095
. PDF.] Reference page.Ponomarenko, A.G. 2021. [Permian Triaplids (Coleoptera, Triaplidae), Yet Another Group of Mesozoic Beetles from the Terminal Permian]. Paleontologicheskii Zhurnal 2021(6): 63–75. (in Russian). [English translation: Paleontological Journal 55(6): 650–661. DOI: 10.1134/S0031030121050130
.] PDF. Reference page.Ponomarenko, A.G., Yan, E.V., Wang, B. & Zhang, H.-C. 2012. Revision of some early Mesozoic beetles from China. Acta Palaeontologica Sinica 51(4): 475–490. Abstract. PDF. Reference page.
Prokin, A.A. & Bashkuev, A.S. 2020 (Online) 2021 (Print). Trialarva coburgensis gen. et sp. nov., a remarkable fossil holometabolan larva (Insecta: Coleoptera) from the Triassic of Germany. PalZ 95: 55–60. DOI: 10.1007/s12542-020-00527-6
. ResearchGate
. Reference page.Qvarnström, M., Fikáček, M., Wernström, J.V., Huld, S., Beutel, R.G., Arriaga-Varela, E., Ahlberg, P.E. & Niedźwiedzki, G. 2021. Exceptionally preserved beetles in a Triassic coprolite of putative dinosauriform origin. Current Biology 31(15): 3374–3381.e5. DOI: 10.1016/j.cub.2021.05.015
. Reference page.Yan, E.V., Beutel, R.G. & Ponomarenko, A.G. 2017 (online) [2018 (print)]. †Peltosynidae, a new beetle family from the Middle–Late Triassic of Kyrgyzstan: its affinities with Polyphaga (Insecta, Coleoptera) and the groundplan of this megadiverse suborder. Journal of Systematic Palaeontology 16(6): 515–530. DOI: 10.1080/14772019.2017.1313789
. ResearchGate
. Reference page.Yan, E.V., Beutel, R.G. & Ponomarenko, A.G. 2017. Ademosynidae (Insecta: Coleoptera): A new concept for a coleopteran key taxon and its phylogenetic affinities to the extant suborders. Palaeontologia Electronica 20.2.31A: 1–22. DOI: 10.26879/739
. Reference page.Andújar, C. & Grebennikov, V.V. 2021. Endogean beetles (Coleoptera) of Madagascar: deep soil sampling and illustrated overview. Zootaxa 4963(2): 317–344. DOI: 10.11646/zootaxa.4963.2.4
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Coleoptera - Australian Faunal Directory
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Beetles and coleopterologists Russian site with English version, with information about biology, systematics and paleontology of beetles
Coleoptera - Coleoptera-atlas (Beautiful Pictures of coleoptera - picture Gallery (over 4800 records)
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