Llorenç Sáez - Profile on Academia.edu (original) (raw)

Papers by Llorenç Sáez

Mediterranean botany, Apr 12, 2021

Potentilla asturica, an Iberian endemic species included within sect. Recta, is typified.

Orsis, organismes i sistemes: revista de botànica, zoologia i ecologia, 2016

This paper deals with 67 taxa, 25 of which are new to the Balearic Islands flora:

Willdenowia, Sep 1, 2015

BioOne Complete (complete.BioOne.org) is a full-text database of 200 subscribed and open-access t... more BioOne Complete (complete.BioOne.org) is a full-text database of 200 subscribed and open-access titles in the biological, ecological, and environmental sciences published by nonprofit societies, associations, museums, institutions, and presses.

International Journal of Plant Sciences, Mar 1, 2009

The genus Helichrysum has 500-600 species in Africa, Madagascar, the Mediterranean basin, Macaron... more The genus Helichrysum has 500-600 species in Africa, Madagascar, the Mediterranean basin, Macaronesia, western and central Asia, and India. The aims of this study are to confirm the exclusion of Castroviejoa from Helichrysum, to elucidate the origin of the Macaronesian endemics, to confirm the monophyly of the Mediterranean-Asiatic group, and, within this group, to contrast molecular data with sectional classification and to provide insight into trends in character evolution and biogeography. To address these issues, ITS and ETS sequences were used in phylogenetic analyses. Results show that Castroviejoa is excluded from Helichrysum. The phylogeny obtained suggests a southern African origin for Helichrysum and several subsequent migrations throughout the African continent. The species from the Canary Islands do not form a monophyletic clade, suggesting that at least two independent colonization events occurred. Helichrysum makranicum, from Oman and southern Iran, is the closest relative of a clade consisting of two endemics of the Canary Islands, providing new evidence of the phytogeographical links between Macaronesia and eastern Africa-southern Arabia. Helichrysum gossypinum, the third endemic of the Canary Islands, is the sister species of a monophyletic Mediterranean and Asiatic group, with the four endemics of Madeira sister to all of them. Within the Mediterranean-Asiatic clade, sect. Stoechadina is monophyletic, but sect. Helichrysum and sect. Virginea are not. Once in the Mediterranean region, the genus is hypothesized to have diversified and expanded to the east up to western and central Asia and to have suffered a reduction in woodiness.

Taxon, Feb 7, 2017

Cymbalaria comprises ten species and six subspecies growing in rocky habitats in the Mediterranea... more Cymbalaria comprises ten species and six subspecies growing in rocky habitats in the Mediterranean Basin. Several features, such as the genus' highly fragmented distribution as well as noticeable ecological differentiation between partially sympatric species and presence of ploidy barriers between species suggest the involvement of different speciation types in its evolution. The aims of this study were to test the monophyly of Cymbalaria and to reconstruct infrageneric phylogenetic relationships, to infer the genus' biogeographic history by estimating divergence times and ancestral distribution areas of lineages, and to disentangle the role of different speciation types. To address these issues, we constructed a phylogeny with a complete taxon sampling based on ITS, 3'ETS, ndhF and rpl32-trnL sequences. We used the nuclear ribosomal DNA data to produce a time-calibrated phylogeny, which served as basis for estimating ploidy level evolution and biogeographic history. Cymbalaria was resolved as monophyletic. The genus originated ca. 4 Ma and three lineages segregated rapidly, one comprising solely C. microcalyx subsp. microcalyx and the other two corresponding to western and central-eastern species, respectively. The main diversification events occurred after the onset of the Mediterranean climate and during Pleistocene climate oscillations. Both founder-event speciation linked to longdistance dispersal events and sympatric speciation were supported by the biogeographic analyses. In addition, at least two polyploid speciation events were inferred. Finally, conflicts between current taxonomy and the phylogeny at the species and subspecies level clearly show the need of more detailed integrative taxonomic studies. Keywords Ancestral-area estimation, cpDNA, founder-event speciation, longdistance dispersal, molecular dating, nrDNA C. aequitriloba 5 C. muralis subsp. muralis 1 C. muralis subsp. muralis 2 C. muelleri 2 C. hepaticifolia 2 C. aequitriloba 6 C. muralis subsp. muralis 4 C. glutinosa subsp. glutinosa 2 C. muelleri 1 C. microcalyx subsp. dodekanesi C. aequitriloba 4 C. hepaticifolia 1 C. aequitriloba 7 C. microcalyx subsp. minor C. pallida 2 C. aequitriloba 1 C. aequitriloba 2 C. glutinosa subsp. glutinosa 1 C. pallida 3 C. muralis subsp. visianii 1 C. pubescens 4 C. glutinosa subsp. brevicalcarata C. muralis subsp. visianii 2 C. longipes 2 C. pubescens 2 C. microcalyx subsp. ebelii C. muralis subsp. visianii 4 C. pallida 1 C. microcalyx subsp. acutiloba C. pubescens 1 C. muralis subsp. muralis 3 C. muralis subsp. visianii 3 C. pubescens 3 C. fragilis 1 C. microcalyx subsp. microcalyx Epixiphium wislizeni C. aequitriloba 3 Asarina procumbens C. longipes 1 C. fragilis 2

En total s'han analitzat 1408 tàxons, 983 dels quals són espècies oròfiles i 317 són, a més, d'al... more En total s'han analitzat 1408 tàxons, 983 dels quals són espècies oròfiles i 317 són, a més, d'alta muntanya. El 9,02% d'espècies són al•lòctones introduïdes, menys de la meitat del registrat a Catalunya; i el 4,63% són espècies raríssimes al país.

Plant Systematics and Evolution, Feb 4, 2021

The eastern Mediterranean basin hosts a remarkably high plant diversity. Historical connections b... more The eastern Mediterranean basin hosts a remarkably high plant diversity. Historical connections between currently isolated areas across the Aegean region and long-distance dispersal events have been invoked to explain current distribution patterns of species. According to most recent treatments, at least two Cymbalaria species occur in this area, Cymbalaria microcalyx and C. longipes. The former comprises several intraspecific taxa, treated at different ranks by different authors based on morphological data, evidencing the need of a taxonomic revision. Additionally, some populations of C. microcalyx show exclusive morphological characters that do not match any described taxon. Here, we aim to shed light on the systematics of eastern Mediterranean Cymbalaria and to propose a classification informed by various sources of evidence. We performed molecular phylogenetic analyses using ITS, 3'ETS, ndhF and rpl32-trnL sequences and estimated the ploidy level of some taxa performing relative genome size measures. Molecular data combined with morphology support the division of traditionally delimited C. microcalyx into C. acutiloba, C. microcalyx and C. minor, corresponding to well-delimited nrDNA lineages. Furthermore, we propose to combine C. microcalyx subsp. paradoxa at the species level. A group of specimens previously thought to belong to Cymbalaria microcalyx constitute a well-defined phylogenetic and morphological entity and are described here as a new species, Cymbalaria spetae. Cymbalaria longipes is non-monophyletic, but characterized by being glabrous and diploid, unlike other eastern species. The nrDNA data suggest at least two dispersals from the mainland to the Aegean Islands, potentially facilitated by marine regressions.

Orsis, organismes i sistemes: revista de botànica, zoologia i ecologia, 2001

Presentem dades sobre l'estatus de diversos endemismes de Catalunya (NE de la península Ibèrica) ... more Presentem dades sobre l'estatus de diversos endemismes de Catalunya (NE de la península Ibèrica) o de plantes rares amb poblacions isolades en aquesta zona. El seu estatus ha estat avaluat aplicant els criteris de la Llista Vermella de la IUCN (2001). Per als tàxons endèmics del territori estudiat, la catalogació proposada és:

Collectanea Botanica, Apr 20, 2020

MicroMeria rodriguezii (Lamiaceae) in the iberian peninsuLar fLora.-Micromeria rodriguezii is an ... more MicroMeria rodriguezii (Lamiaceae) in the iberian peninsuLar fLora.-Micromeria rodriguezii is an endemic species from the Balearic Islands (Western Mediterranean basin). A population of this species has been found in NE Spain (Castellón province), growing in the margins of a forest track. A study of the plant morphology of this Iberian population is provided. Its conservation status in the Valencian Community is discussed and a management and conservation plan is suggested to be drawn up to ensure its preservation.

When on board of H. M. S 'Beagle', I was much struck with certain facts in the distribution of in... more When on board of H. M. S 'Beagle', I was much struck with certain facts in the distribution of inhabitants of South America, and in the geological relations of the present to the past inhabitants of that continent. These facts seemed to me to throw some light on the origin of the species ̶ that mistery of misterys" Als barrancs, que ara sovintejaven, s'hi escampaven gleves denses de silene brodades de floretes rosades i ornades de l'exuberància de les semprevives, mentre que a les concavitats petites treien el cap les últimes anemones, els ranuncles de les geleres, migrats clavells i centenars d'altres flors diminutes de noms estranys i colors vivíssims. W. Bonatti, 1999 (traducció de S. Pohl) "When on board of H. M. S 'Beagle', I was much struck with certain facts in the distribution of inhabitants of South America, and in the geological relations of the present to the past inhabitants of that continent. These facts seemed to me to throw some light on the origin of the species ̶ that mistery of misterys" Als barrancs, que ara sovintejaven, s'hi escampaven gleves denses de silene brodades de floretes rosades i ornades de l'exuberància de les semprevives, mentre que a les concavitats petites treien el cap les últimes anemones, els ranuncles de les geleres, migrats clavells i centenars d'altres flors diminutes de noms estranys i colors vivíssims. W. Bonatti, 1999 (traducció de S. Pohl) Una tesi sol trobar-se envoltada d'una certa aura de transcendència en l'imaginari col•lectiu. Per fer-ne justícia, aquí em trobo, escrivint la part que més persones llegiran en un tren de la línia R3. Abans de res, però, ho dic o rebento: m'ho he passat molt bé fent aquesta tesi. En aquell penúltim any de carrera, vaig ser víctima de l'entusiasme de la Mercè i el Llorenç per la flora i la distribució dels endemismes. Els vostres estils tan diferents em van inspirar en aquelles memorables jornades de camp en què se'm descobria el tresor que tenia sota els peus. Unes primeres col•laboracions estudiant el gènere Helichrysum van relligar la fascinació per la biodiversitat amb una creixent inquietud en l'Evolució. D'aquesta manera vaig conèixer l'Institut Botànic de Barcelona i la Núria, que des de llavors es va atrevir a pensar que alguna cosa se'n podria fer, de mi. La tesi va anar venint com una seqüència natural, i per una sèrie de caramboles les cimbalàries em van, en certa manera, trobar i convertir-se en les protagonistes d'aquesta tesi. Gràcies, directors, per acollir-me, deixar-me provar coses noves i equivocar-me. Em dec també a tants altres botànics que han anat marcant els meus inicis. Al Ramon Pérez, per dirigir-me a en Llorenç quan et vaig comentar les meves inquietuds. A en Pere Fraga i en Jaume Soler, quin plaer anar al camp amb vosaltres! To all the team from the Evolutionary Systematics Lab in Innsbruck, and specially to Peter, for trusting me and your honesty. The stay in Innsbruck was a milestone in my career. I also thank Pam and Doug Soltis, for being always so positive and open-minded, I learnt a different (and successful!) way of working than we do here. A tot l'equip

Sierra de Alcubierre is the main orographic nucleus of the Monegros and the one which penetrates ... more Sierra de Alcubierre is the main orographic nucleus of the Monegros and the one which penetrates farthest into the most arid part of the Ebro valley. It constitutes a highly interesting biogeographical meeting place: the sharp contrast between the northem and southem slopes gives rise to widely differentiated microclimates offering a broad range of habitats for taxa and plant communities of very varied geographical origins and ecological conditions. Provisional analysis has been carried out on the basis of the 578 taxa so far recognized in this territory in order to establish the percentage of the main chorologic Elements and Subelements. The fioristic background is clearly Mediterranean (65%). Ofparticular significance, in the arid environments where they are located, are the eurosiberian species (8,2%): Atropa belladona, Filipendula vulgaris, f.ex.; also the submediterranean species (6%) are present: examples reported for the first time include Buxus sempervirens, Amelancher ovalis, Polygonatum odoratum, Lonicera xylosteum and Viola willkommii. Outstanding among endemics (8%) are certain territorially endemic taxa such as Allium pardoi, Nepeta beltranii, Vale- rianella multidentata and Reseda lutea subsp. vivantii. The discovery of noteworthy plants endemic to the Iberian peninsula, such as Arrhenaterum album, Bifora radians, Cirsium ferox, Senecio lagascanus, also deserves mention. A new taxon is described: Viola rupestris susbp. orioli-bolosii Molero, L. Sáez & Villar. In terms of vegetation, the phytosociological method was used to identify the communities which provide the best definition of the Plant Landscape of the Range. Of special interest is the study of continental holm-oak forests, Quercetum rotun- difoliae, of which two new subassociations are described: senecietosum lagascani Molero, Sáez & Villar and viburnetosum tini Molero, Sáez & Villar. Impoverished gall-oak groves -Violo-Quercetumfagineae subass. viburnetosum tini Molero, Sáez Trabajo financiado con cargo al proyecto "Recursos fitogenéticos de Aragón", desarrollado por

The precise generic delimitation of Aliella and Phagnalon, and their closest relatives within the... more The precise generic delimitation of Aliella and Phagnalon, and their closest relatives within the Gnaphalieae are discussed in this review. Among the main results obtained, we have found that the genera Aliella and Phagnalon are nested within the "Relhania clade" and Anisothrix, Athrixia and Pentatrichia are their closest relatives. Macowania is also part of the "Relhania clade", whereas the subtribal affinities of Philyrophyllum lie within the "crown radiation clade". The monophyly of Aliella and Phagnalon is not supported statistically. In addition, Aliella appears to be paraphylethic in most of the analyses performed. The resulting phylogeny suggests an African origin for the ancestor of Aliella and Phagnalon and identifies three main clades within Phagnalon that constitute the following natural groups on a geographic basis: (1) the Irano-Turanian clade; (2) the Mediterranean-Macaronesian clade; and (3) the Yemen-Ethiopian

FIGURE 11. Female florets. A in Evaluation of the taxonomic status of the genus Aliella (Compositae, Gnaphalieae): a recircumscription of the genus Phagnalon

FIGURE 11. Female florets. A. Aliella ballii (Sáez 6191, Sáez pers. herb.) B. Phagnalon latifoliu... more FIGURE 11. Female florets. A. Aliella ballii (Sáez 6191, Sáez pers. herb.) B. Phagnalon latifolium (Charpin et al. MAR 206, W). C. Phagnalon stenolepis (Collenette 9135, E), (A–C) scale bar = 1 mm. Waxy cushions on the corolla lobes. D. Phagnalon carolipaui (Sauvage 12374, MA), scale bar = 20 µm.

New data on the distribution of Limonium catalaunicum (Plumbaginaceae) in Catalonia (north-easter... more New data on the distribution of Limonium catalaunicum (Plumbaginaceae) in Catalonia (north-eastern Iberian Peninsula). It has been difficult to determine the distribution of Limonium catalanicum because, for many years, this species has been confused with other species of the genus. In this article we provide new data about this species and we detail the 7 populations known at the moment. In as much, we report 6 populations that might be extinguished or that they haven't been found recently, so we contribute to the clarification of its current distribution. Finally, we review the threat level of this species that should remain as critical despite the increase of known populations.

Taraxacum majoricense(Asteraceae), a New Species from the Balearic Islands, Spain

Annales Botanici Fennici, 2016

Taraxacum majoricense A. Galán & L. Sáez sp. nova collected from Mallorca in the Balearic Isl... more Taraxacum majoricense A. Galán & L. Sáez sp. nova collected from Mallorca in the Balearic Islands is described and illustrated. It is characterized by rough leaves with wide lobes, and by violet achenes. A key to all species of Taraxacum known from the Balearic Islands is provided. Taraxacum tarraconense Sennen is neotypified and newly described. Taraxacum cacuminatum, T. catalanum, T. marklundii and T. tarraconense are new records for the Balearic Islands.

Type designation of the Lamarck’s name Buxus balearica (Buxaceae)

Phytotaxa, Aug 29, 2019

The Lamarck’s name Buxus balearica is discussed and neotypified from a specimen preserved at MA c... more The Lamarck’s name Buxus balearica is discussed and neotypified from a specimen preserved at MA collected in Mallorca (Balearic Islands, Spain). In addition, the lectotypes of B. longifolia and B. balearica f. granatensis, currently treated as two heterotypic synonyms of B. balearica, are designated from original material preserved at G-BOIS and MA, respectively

Orsis, Nov 22, 2017

This compilation includes nomenclatural and taxonomic updates to the checklist of vascular plants... more This compilation includes nomenclatural and taxonomic updates to the checklist of vascular plants described from the Balearic Islands. This update includes plants described between 2008 and 2016, and contains corrections to other entries included in previous works.