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Papers by Charles Bartlett

Research paper thumbnail of A new species of planthopper in the genus Colpoptera (Hemiptera: Fulgoroidea: Nogodinidae) from the Northeastern coast of Jamaica

Research paper thumbnail of A new species of planthopper in the genus Paraphenice (Hemiptera: Derbidae: Otiocerinae) from palms in eastern Madagascar

Research paper thumbnail of A new species of planthopper in the genus Anotia (Hemiptera: Auchenorrhyncha: Derbidae) from the Los Angeles cloud forest in Costa Rica

Research paper thumbnail of Summary and Discussion

Research paper thumbnail of Figures

Research paper thumbnail of References Cited

Research paper thumbnail of Systematic Treatment

Research paper thumbnail of Phylogenetic Analysis of <i>Pissonotus</i&gt

Research paper thumbnail of Appendices

Research paper thumbnail of <p><strong>A new species of planthopper belonging to the genus <em>Oecleus</em> Stål, 1862 (Hemiptera: Fulgoroidea: Cixiidae) from coconut palm (<em>Cocos</em> <em>nucifera</em> L) in Jamaica</strong></p&gt

Research paper thumbnail of A new species of planthopper in the genus Haplaxius from Osa Peninsula in Costa Rica (Hemiptera: Fulgoroidea: Cixiidae)

Research paper thumbnail of A new species of Oecleus (Hemiptera: Auchenorrhyncha: Fulgoroidea: Cixiidae) from the Caribbean coast of Costa Rica and a reassessment of the generic status of Nymphomyndus

Zootaxa

A new species of cixiid planthopper in the genus Oecleus Stål is described from Tortuguero, Limón... more A new species of cixiid planthopper in the genus Oecleus Stål is described from Tortuguero, Limón Province, Costa Rica. This is the first Oecleus species reported from Costa Rica. The new species, Oecleus dormido sp. n., was collected sweeping grassy edge habitat near the Caribbean coast. Sequence data for COI, 18S, and H3 was generated for phylogenetic comparison of the new species with other available Oecleus species and New World Oecleini to test genus-level placement and provide a preliminary examination of the phylogenetic relationships among New World genera of Oecleini. The results indicate that Oecleus, Haplaxius and Myxia are monophyletic, at least among the included taxa, the genera (Myxia+((Nymphocixia+Nymphomyndus)+Haplaxius) may form a clade apart from Oecleus. The monobasic genera Nymphocixia+Nymphomyndus are closely allied, with a difference of 0.98% for 18S, compared to an average of 2.2% among other included oecleine genera. In view of the limited morphological and ...

Research paper thumbnail of First New World Record ofParadelphacodes Paludosus(Flor 1861) (Hemiptera: Fulgoroidea : Delphacidae) in Alaska

Entomological News, Apr 1, 2015

Research paper thumbnail of Systematics of Hadropygos n.g., Metadelphax Wagner and New World Toya Distant (Hemiptera: Delphacidae)

Transactions of The American Entomological Society, Sep 1, 2007

New World members of the genera Toya Distant and Metadelphax Wagner are here revised and the genu... more New World members of the genera Toya Distant and Metadelphax Wagner are here revised and the genus Hadropygos n. g. described from South America. Generic limits are discussed, with particular comparison to Syndelphax Fennah. A key to treated genera and species is provided. The type species of Toya, T. attenuata Distant from Sri Lanka, is illustrated and discussed. Among the New World Toya, T. boxi (Muir) and T. venilia (Fennah) are retained in the genus; two species are transferred to Toya from Delphacodes Fieber, Toya idonea (Beamer) new comb. and T. nigra (Crawford) new comb., with Delphacodes axonopi Fennah placed as a new junior synonym of T. nigra. Three new species of New World Toya are described (T. goliai n. sp., T. dietrichi n. sp., and T. recurva n. sp.). Toya iaxartes (Fennah), described from a single specimen from St. Lucia, is here considered nomen dubium as it appears to have been described from a malformed or parasitized specimen. Metadelphax, restored as a genus by Ding (2006), consists of five species: M. propinqua (Fieber) (type species), M. argentinensis new comb., here transferred from Toya, M. pero, here transferred from Syndelphax, M. wetmorei (Muir and Giffard) new comb., here transferred from Delphacodes, plus M. dentata n. sp. Metadelphax bridwelli (Muir), transferred from Toya by Ding (2006), is returned to Toya. Hadropygos n. g., with the single species, H. rhombos n. sp., from the Neotropics bears comparable features to Toya and Metadelphax. Lectotype designations are made for Toya attenuata Distant and Delphax propinqua Fieber. Collectively, 3 genera are treated: Hadropygos n. g., with a single new species, Metadelphax with 5 species (1 new), and Toya with 7 New World species (3 new), excluding T. iaxartes as a nomen dubium.

Research paper thumbnail of Planthopper (Hemiptera: Fulgoroidea) Diversity in the Great Smoky Mountains National Park

Transactions of The American Entomological Society, Sep 1, 2006

Planthoppers from Malaise traps (22 traps operated continuously over 3 years at 11 sites) were in... more Planthoppers from Malaise traps (22 traps operated continuously over 3 years at 11 sites) were investigated as a component of an ongoing all-taxon biotic inventory of the Great Smoky Mountains National Park. The Malaise samples contained 2,195 specimens representing 55 species from 7 families. Elevation patterns and seasonality of planthoppers from the Malaise traps are also presented. Based on Malaise trap data, species accumulation curves and 9 estimators of species richness anticipated 57-81 species, a higher number than predicted in our previous study (Bartlett & Bowman 2004), but lower than the cumulative species list, currently including 97 species in 10 families based on 6,860 specimens. None of the data sets based on single sample methods produced estimates of species richness that were greater than the observed diversity from the compiled species list. Species richness estimates between 95-124 were obtained by combining the Malaise trap sample data with Bartlett and Bowman's (2004) sweep sample data and additional data from 2006, although two estimators still remained below the observed richness of 97 species for the Park. Implications of these findings are discussed.

Research paper thumbnail of Planthoppers of Delaware (Hemiptera, Fulgoroidea), excluding Delphacidae, with species Incidence from adjacent States

ZooKeys, Feb 25, 2011

Th e number of species of planthoppers (excluding Delphacidae) known from Delaware is updated fro... more Th e number of species of planthoppers (excluding Delphacidae) known from Delaware is updated from 7 (in 4 families) to 62 species (in 9 families). Specimen abundance is tallied by county and seasonally by two week intervals. Th e Chao1 abundance estimator suggests that the true fauna may be 74 species, although species incidence tallied from adjacent states (MD, NJ, PA and DC) suggests that a total fauna of approximately 100 species may be possible. An artifi cial key is presented to genus and select species with photos of most included taxa.

Research paper thumbnail of The Diversity of the True Hoppers (Hemiptera: Auchenorrhyncha)

Insect Biodiversity, 2018

Research paper thumbnail of On the Generic Placement and Synonymy for Delphacodes guianensis Muir 1919 (Hemiptera: Fulgoroidea: Delphacidae)

Entomological News, May 25, 2021

ABSTRACT: The holotype of Delphacodes guianensis Muir, a delphacid planthopper (Hemiptera: Fulgor... more ABSTRACT: The holotype of Delphacodes guianensis Muir, a delphacid planthopper (Hemiptera: Fulgoroidea: Delphacidae) currently nominally in the polyphyletic genus Delphacodes Fieber, was examined and found to bear lateral teeth on the hind basitarsus, a synapomorphy of the genus Nilaparvata Distant. We determined that Delphacodes guianensis is a senior synonym of Nilaparvata serrata Caldwell. Here we move this species into Nilaparvata to provide the new combination Nilaparvata guianensis (Muir) and give Nilaparvata serrata as a new junior synonym.

Research paper thumbnail of An Unusual Morphotype of Caenodelphax teapae (Fowler) (Hemiptera: Fulgoroidea: Delphacidae)

Entomological News, 2015

Caenodelphax teapae (Fowler) is a common, widespread Neotropical species. This species is potenti... more Caenodelphax teapae (Fowler) is a common, widespread Neotropical species. This species is potentially pestiferous as it has been reported on carrots, sugar cane, kidney beans and other crops, and is a vector of Urochloa hoja blanca tenuivirus in plantain signalgrass ( Urochloa plantaginea (Link) R. Webster) (Lapierra and Signoret 2004, Wilson 2005, Kennedy and Bartlett 2014). The species was first described from Teapa, Mexico in 1905 (in Liburnia Stal), and subsequently established as the type species of Caenodelphax Fennah, 1965 (by original designation). Caenodelphax was recently revised and is now monotypic (Kennedy and Bartlett 2014). Here we report the unusual retention of juvenile features (i.e., paedomorphosis) in adult C. teapae . Ceanodelphax teapae exhibits a predictable level of geographical variation in coloration and morphology. Individuals are usually macropterous and between approximately 2.0-3.5 mm in length, dark brown to black in body color, with concolorous carinae and paler (yellow) antennae, beak and legs (Figs. 1A-C). Females are usually larger and paler than males. Brachypters are uncommon (and usually female) and much paler than macropters, but bear a comparable color pattern.

Research paper thumbnail of New State Records of Treehoppers (Hemiptera: Membracidae) for Delaware, Maryland, Mississippi, Pennsylvania, And West Virginia

Proceedings of The Entomological Society of Washington, Jul 1, 2009

Research paper thumbnail of A new species of planthopper in the genus Colpoptera (Hemiptera: Fulgoroidea: Nogodinidae) from the Northeastern coast of Jamaica

Research paper thumbnail of A new species of planthopper in the genus Paraphenice (Hemiptera: Derbidae: Otiocerinae) from palms in eastern Madagascar

Research paper thumbnail of A new species of planthopper in the genus Anotia (Hemiptera: Auchenorrhyncha: Derbidae) from the Los Angeles cloud forest in Costa Rica

Research paper thumbnail of Summary and Discussion

Research paper thumbnail of Figures

Research paper thumbnail of References Cited

Research paper thumbnail of Systematic Treatment

Research paper thumbnail of Phylogenetic Analysis of <i>Pissonotus</i&gt

Research paper thumbnail of Appendices

Research paper thumbnail of <p><strong>A new species of planthopper belonging to the genus <em>Oecleus</em> Stål, 1862 (Hemiptera: Fulgoroidea: Cixiidae) from coconut palm (<em>Cocos</em> <em>nucifera</em> L) in Jamaica</strong></p&gt

Research paper thumbnail of A new species of planthopper in the genus Haplaxius from Osa Peninsula in Costa Rica (Hemiptera: Fulgoroidea: Cixiidae)

Research paper thumbnail of A new species of Oecleus (Hemiptera: Auchenorrhyncha: Fulgoroidea: Cixiidae) from the Caribbean coast of Costa Rica and a reassessment of the generic status of Nymphomyndus

Zootaxa

A new species of cixiid planthopper in the genus Oecleus Stål is described from Tortuguero, Limón... more A new species of cixiid planthopper in the genus Oecleus Stål is described from Tortuguero, Limón Province, Costa Rica. This is the first Oecleus species reported from Costa Rica. The new species, Oecleus dormido sp. n., was collected sweeping grassy edge habitat near the Caribbean coast. Sequence data for COI, 18S, and H3 was generated for phylogenetic comparison of the new species with other available Oecleus species and New World Oecleini to test genus-level placement and provide a preliminary examination of the phylogenetic relationships among New World genera of Oecleini. The results indicate that Oecleus, Haplaxius and Myxia are monophyletic, at least among the included taxa, the genera (Myxia+((Nymphocixia+Nymphomyndus)+Haplaxius) may form a clade apart from Oecleus. The monobasic genera Nymphocixia+Nymphomyndus are closely allied, with a difference of 0.98% for 18S, compared to an average of 2.2% among other included oecleine genera. In view of the limited morphological and ...

Research paper thumbnail of First New World Record ofParadelphacodes Paludosus(Flor 1861) (Hemiptera: Fulgoroidea : Delphacidae) in Alaska

Entomological News, Apr 1, 2015

Research paper thumbnail of Systematics of Hadropygos n.g., Metadelphax Wagner and New World Toya Distant (Hemiptera: Delphacidae)

Transactions of The American Entomological Society, Sep 1, 2007

New World members of the genera Toya Distant and Metadelphax Wagner are here revised and the genu... more New World members of the genera Toya Distant and Metadelphax Wagner are here revised and the genus Hadropygos n. g. described from South America. Generic limits are discussed, with particular comparison to Syndelphax Fennah. A key to treated genera and species is provided. The type species of Toya, T. attenuata Distant from Sri Lanka, is illustrated and discussed. Among the New World Toya, T. boxi (Muir) and T. venilia (Fennah) are retained in the genus; two species are transferred to Toya from Delphacodes Fieber, Toya idonea (Beamer) new comb. and T. nigra (Crawford) new comb., with Delphacodes axonopi Fennah placed as a new junior synonym of T. nigra. Three new species of New World Toya are described (T. goliai n. sp., T. dietrichi n. sp., and T. recurva n. sp.). Toya iaxartes (Fennah), described from a single specimen from St. Lucia, is here considered nomen dubium as it appears to have been described from a malformed or parasitized specimen. Metadelphax, restored as a genus by Ding (2006), consists of five species: M. propinqua (Fieber) (type species), M. argentinensis new comb., here transferred from Toya, M. pero, here transferred from Syndelphax, M. wetmorei (Muir and Giffard) new comb., here transferred from Delphacodes, plus M. dentata n. sp. Metadelphax bridwelli (Muir), transferred from Toya by Ding (2006), is returned to Toya. Hadropygos n. g., with the single species, H. rhombos n. sp., from the Neotropics bears comparable features to Toya and Metadelphax. Lectotype designations are made for Toya attenuata Distant and Delphax propinqua Fieber. Collectively, 3 genera are treated: Hadropygos n. g., with a single new species, Metadelphax with 5 species (1 new), and Toya with 7 New World species (3 new), excluding T. iaxartes as a nomen dubium.

Research paper thumbnail of Planthopper (Hemiptera: Fulgoroidea) Diversity in the Great Smoky Mountains National Park

Transactions of The American Entomological Society, Sep 1, 2006

Planthoppers from Malaise traps (22 traps operated continuously over 3 years at 11 sites) were in... more Planthoppers from Malaise traps (22 traps operated continuously over 3 years at 11 sites) were investigated as a component of an ongoing all-taxon biotic inventory of the Great Smoky Mountains National Park. The Malaise samples contained 2,195 specimens representing 55 species from 7 families. Elevation patterns and seasonality of planthoppers from the Malaise traps are also presented. Based on Malaise trap data, species accumulation curves and 9 estimators of species richness anticipated 57-81 species, a higher number than predicted in our previous study (Bartlett & Bowman 2004), but lower than the cumulative species list, currently including 97 species in 10 families based on 6,860 specimens. None of the data sets based on single sample methods produced estimates of species richness that were greater than the observed diversity from the compiled species list. Species richness estimates between 95-124 were obtained by combining the Malaise trap sample data with Bartlett and Bowman's (2004) sweep sample data and additional data from 2006, although two estimators still remained below the observed richness of 97 species for the Park. Implications of these findings are discussed.

Research paper thumbnail of Planthoppers of Delaware (Hemiptera, Fulgoroidea), excluding Delphacidae, with species Incidence from adjacent States

ZooKeys, Feb 25, 2011

Th e number of species of planthoppers (excluding Delphacidae) known from Delaware is updated fro... more Th e number of species of planthoppers (excluding Delphacidae) known from Delaware is updated from 7 (in 4 families) to 62 species (in 9 families). Specimen abundance is tallied by county and seasonally by two week intervals. Th e Chao1 abundance estimator suggests that the true fauna may be 74 species, although species incidence tallied from adjacent states (MD, NJ, PA and DC) suggests that a total fauna of approximately 100 species may be possible. An artifi cial key is presented to genus and select species with photos of most included taxa.

Research paper thumbnail of The Diversity of the True Hoppers (Hemiptera: Auchenorrhyncha)

Insect Biodiversity, 2018

Research paper thumbnail of On the Generic Placement and Synonymy for Delphacodes guianensis Muir 1919 (Hemiptera: Fulgoroidea: Delphacidae)

Entomological News, May 25, 2021

ABSTRACT: The holotype of Delphacodes guianensis Muir, a delphacid planthopper (Hemiptera: Fulgor... more ABSTRACT: The holotype of Delphacodes guianensis Muir, a delphacid planthopper (Hemiptera: Fulgoroidea: Delphacidae) currently nominally in the polyphyletic genus Delphacodes Fieber, was examined and found to bear lateral teeth on the hind basitarsus, a synapomorphy of the genus Nilaparvata Distant. We determined that Delphacodes guianensis is a senior synonym of Nilaparvata serrata Caldwell. Here we move this species into Nilaparvata to provide the new combination Nilaparvata guianensis (Muir) and give Nilaparvata serrata as a new junior synonym.

Research paper thumbnail of An Unusual Morphotype of Caenodelphax teapae (Fowler) (Hemiptera: Fulgoroidea: Delphacidae)

Entomological News, 2015

Caenodelphax teapae (Fowler) is a common, widespread Neotropical species. This species is potenti... more Caenodelphax teapae (Fowler) is a common, widespread Neotropical species. This species is potentially pestiferous as it has been reported on carrots, sugar cane, kidney beans and other crops, and is a vector of Urochloa hoja blanca tenuivirus in plantain signalgrass ( Urochloa plantaginea (Link) R. Webster) (Lapierra and Signoret 2004, Wilson 2005, Kennedy and Bartlett 2014). The species was first described from Teapa, Mexico in 1905 (in Liburnia Stal), and subsequently established as the type species of Caenodelphax Fennah, 1965 (by original designation). Caenodelphax was recently revised and is now monotypic (Kennedy and Bartlett 2014). Here we report the unusual retention of juvenile features (i.e., paedomorphosis) in adult C. teapae . Ceanodelphax teapae exhibits a predictable level of geographical variation in coloration and morphology. Individuals are usually macropterous and between approximately 2.0-3.5 mm in length, dark brown to black in body color, with concolorous carinae and paler (yellow) antennae, beak and legs (Figs. 1A-C). Females are usually larger and paler than males. Brachypters are uncommon (and usually female) and much paler than macropters, but bear a comparable color pattern.

Research paper thumbnail of New State Records of Treehoppers (Hemiptera: Membracidae) for Delaware, Maryland, Mississippi, Pennsylvania, And West Virginia

Proceedings of The Entomological Society of Washington, Jul 1, 2009