José E García Raso | Universidad de Málaga (original) (raw)
Papers by José E García Raso
Bulletin of Marine Science, 1996
During the Balgim-84 Expedition, carried out from both sides of the Straits of Gibraltar (Atlanti... more During the Balgim-84 Expedition, carried out from both sides of the Straits of Gibraltar (Atlantic Ocean: Gulf of Cadiz and Mediterranean Sea: Alboran Sea), in depths between 137 and 2,142 m, 3,559 specimens of decapods belonging to 122 species were captured. Many of these species are rare, of biogeographical interest and represent new records for the area. For some species, anatomical details are given. A specimen of Plesionika may be a new species. The collection of Calocarides coronatus, Sympagurus acinops, Monodaeus guinotae, Periclimenes kornii, Alpheus talismani, Athanas amazone, Merhippolyte ancitrosta, Eualus cf. lebourae, Plesionika williamsi, Pontophilus cf. abyssi, Spongicoloides evolutus, Spongicoloides profundus, Odontozona edwardsi, Catapaguroides iris and Strobopagurus gracilipes, among others, represents new records for the Mediterranean Sea, eastern Atlantic, and Iberian waters.
Invertebrate Biology, 2005
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Cahiers De Biologie Marine, 1992
The collect of a male specimen of Micl'ocassiope lIIinol' (Dana) from the southeast of Spain repr... more The collect of a male specimen of Micl'ocassiope lIIinol' (Dana) from the southeast of Spain represe nts the first record of the species for the West Mediterranean Sea. The recent studies of the decapod faLma of the Alboran Sea show that West African Atl antic species are co lonizing thi s Mediterranean area. , Résumé: Le déve loppement récent des recherches sur les Décapodes dans la Mer de Alboran (à l' est de Gibraltar) montre l' extensio n d' espèces ouest-africains en Méditerranée. Microcassiope minol' (Dana) , récoltée dans des cavités d'algues corallinacées, est signalée pour la première fois sur les côtes européennes de la Méditerranée occidentale (Espagne).
Cahiers De Biologie Marine, 1992
The study of the evolution of a Crustacea Decapoda community from calcareous bottoms - strongly r... more The study of the evolution of a Crustacea Decapoda community from calcareous bottoms - strongly related with Posidonia oceanica, developed during 1986-1987. completes the overview of the structure and varia- bility existing in this habitat. Four groups have been stablished, according to the abundance-presence. The cualita- tive composition is similar to that found in other calcareous bottoms from the Southern Spain. So, the six more abundant species (belonging to the first and second groups) are the same. The fluctuations of the general structure and its evolution are controled by the two dominant species, Athanas nitescens and Pisidia longicomis (form. lon- gimana). The development of the Posidonia bed. which increases the sedimentation and protection, is, possibly, the responsible of the cuantitative differences found.
Scientia Marina
The unique colour patterns of the hermit crab species inhabiting the Iberian Peninsula and geogra... more The unique colour patterns of the hermit crab species inhabiting the Iberian Peninsula and geographically close areas are studied based on colour patterns observed in live specimens either in the field or live in the laboratory. Live colour patterns are shown to be useful for differentiating species in the Paguroidea from Iberian waters. Colour information has not been frequently documented in previous studies, mainly because of difficulties in accessing live specimens. Up to 51 species are currently recorded within the study area, 45 of which are included in the present work, focusing on the distinctive colour traits for each species that can be observed in the field. A complete key for identifying the species within the study area based on colour patterns is included. This study is the first attempt to put in place this type of tool oriented to field work. Further studies will be required to complete and update this information, especially for species which are scarce or difficult...
Scientia Marina, 2008
The structure of decapod crustacean assemblages living in shallow waters (5 to 25 m depth) in an ... more The structure of decapod crustacean assemblages living in shallow waters (5 to 25 m depth) in an area of the Alborán Sea (Mediterranean Sea) neighbouring the Strait of Gibraltar was studied. The relatively high richness found in this area is probably due to the diversity of substrata and the confluence of Atlantic and Mediterranean waters. The most abundant species was the hermit crab Diogenes pugilator due to the prevalence of sandy substrate, the shallow depth-range, and the species' life history. The coralligenous bottom showed the highest species richness and diversity values due to the complexity of microhabitats in this type of bottom and probably due to the food flux associated with currents. Conversely, shallow, well calibrated, fine to medium sandy bottoms of 5 m had the lowest values and the maximum densities due to the high abundance of a few well adapted species. The analysis of the different stations showed significant spatial differences according to depth and sedimentary characteristics. Depth is the environmental variable that correlates best with the decapod assemblages, with a particularly significant boundary between 5 m and 15 m. Nevertheless, there was a continuous transition between the assemblages. These results evidence the importance of quantitative studies in differentiating decapod assemblages. Relationships between these assemblages should also be taken into account in coastal management, since altering a substrate could have repercussions for the structure of the communities of neighbouring substrates. No seasonal significant differences were found in the overall analysis of the area, but there were differences between spring and autumn and spring and summer in relation to depth and substrate (crossed analysis). Finally, we present the species which allow us to discriminate the different assemblages according to sediment and depth, as well as the species' contributions.
Scientia Marina, 2002
The present paper presents an overview of the studies on the genus Paguristes from the Caribbean ... more The present paper presents an overview of the studies on the genus Paguristes from the Caribbean Sea and more specifically from Cuban waters. Additionally, the hermit crabs belonging to the genus Paguristes collected during different surveys of the "Instituto de Oceanología de Cuba" on the Cuban coast have been listed. A new species of hermit crab, Paguristes hologuinensis sp. nov., is described and remarks on a new record, Paguristes paraguanensis MacLaughlin and Provenzano, 1974, are given.
nean. Arxius de Miscel·lànea Zoològica, vol. 7: 72–77. New records of Parasquilla ferussaci (Roux... more nean. Arxius de Miscel·lànea Zoològica, vol. 7: 72–77. New records of Parasquilla ferussaci (Roux, 1830) (Crustacea, Stomatopoda) from the Eastern Atlantic and Western Mediterranean. — We report the occurrence of the little known stomatopod Parasquilla ferussaci on the Atlantic and Mediterranean coasts of the Iberian peninsula. Documentation is based on three specimens captured off Isla Cristina (Huelva) in the Gulf of Cadiz, off Fuengirola (Málaga) in the Alboran Sea and off Gavà (Barcelona) in the North–Western Mediterranean. These reports fill the distribution gap between Eastern Central Atlantic reports and previous Mediterranean reports east of the Balearic Islands.
Marine Protected Areas (MPAs) and Key Biodiversity Areas (KBAs) are important tools for preservin... more Marine Protected Areas (MPAs) and Key Biodiversity Areas (KBAs) are important tools for preserving species, habitats and ecological-geological-oceanographical processes. Nowadays, more than 10% of the Alboran Sea is under some protected figure or is on its way to be protected. This chapter presents the current state of the MPAs and KBAs of the Alboran Sea. Most of the habitats included in the Annex I of the Habitats Directive or the Barcelona Convention are represented in MPAs-KBAs of the Alboran Sea. More than 290 threatened species are also occurring in these MPAs-KBAs, but some of them have experienced a strong decline even within them (e.g. Zostera marina, Pinna nobilis, etc.). Although important progress in the protection of the Alboran Sea has been made in the last decades, there is still much to be done for effective conservation of the habitats and species. The main efforts may focus on improving (1) knowledge of the habitats and species, (2) the surveillance and mitigation against current impacts, (3) the coordination between local, national and international agencies and (4) the measures for improving habitats and threatened species populations, among others. A deep change in the socioeconomic model is then needed for preserving the Alboran Sea natural heritage, promoting then a sustainable use of its resources.
Mediterranean Marine Science, 2012
More than 60 marine non-indigenous species (NIS) have been removed from previous lists and 84 spe... more More than 60 marine non-indigenous species (NIS) have been removed from previous lists and 84 species have been added, bringing the total to 986 alien species in the Mediterranean [775 in the eastern Mediterranean (EMED), 249 in the central Mediterranean (CMED), 190 in the Adriatic Sea (ADRIA) and 308 in the western Mediterranean (WMED)]. There were 48 new entries since 2011 which can be interpreted as approximately one new entry every two weeks. The number of alien species continues to increase, by 2-3 species per year for macrophytes, molluscs and polychaetes, 3-4 species per year for crustaceans, and 6 species per year for fish. The dominant group among alien species is molluscs (with 215 species), followed by crustaceans (159) and polychaetes (132). Macrophytes are the leading group of NIS in the ADRIA and the WMED, reaching 26-30% of all aliens, whereas in the EMED they barely constitute 10% of the introductions. In the EMED, molluscs are the most species-rich group, followed b...
Journal of Crustacean Biology
Integrative taxonomy studies have allowed us to clarify some taxonomic problems in cryptic specie... more Integrative taxonomy studies have allowed us to clarify some taxonomic problems in cryptic species within species of InachusWeber, 1795 with two protogastric tubercles found in European waters. Several morphotypes of Inachus phalangium (Fabricius, 1775) are recognized, and a new species is described from the Atlantic coast of the Iberian Peninsula, for which distribution data are provided. Furthermore, two Inachus cf. thoracicus specimens with a unique sternal morphology were collected from Málaga Spain, but their preservation in formaldehyde prevented molecular analyses. Specimens of I. guentheri (Miers, 1879) have been re-examined and the validity of previous reports from tropical and subtropical Atlantic waters is discussed. These results allow us to separate and clarify the status of species within this group. An identification key for Inachus species with two protogastric tubercles is also provided.
Información del artículo Aportaciones al conocimiento de la fauna y flora litoral de la isla de A... more Información del artículo Aportaciones al conocimiento de la fauna y flora litoral de la isla de Alborán (España).
ABSTRACT The state-of-art on alien species in the Mediterranean Sea is presented, making distinct... more ABSTRACT The state-of-art on alien species in the Mediterranean Sea is presented, making distinctions among the four subregions defined in the EU Marine Strategy Framework Directive: (i) the Western Mediterranean Sea (WMED); (ii) the Central Mediterranean Sea (CMED); (iii) the Adriatic Sea (ADRIA); and (iv) the Eastern Mediterranean Sea (EMED). The updated checklist (December 2010) of marine alien species within each subregion, along with their acclimatization status and origin, is provided. A total of 955 alien species is known in the Mediterranean, the vast majority of them having being introduced in the EMED (718), less in the WMED (328) and CMED (267) and least in the Adriatic (171). Of these, 535 species (56%) are established in at least one area. Despite the collective effort of experts who attempted in this work, the number of introduced species remains probably underestimated. Excluding microalgae, for which knowledge is still insufficient, aliens have increased the total species richness of the Mediterranean Sea by 5.9%. This figure should not be directly read as an indication of higher biodiversity, as spreading of so many aliens within the basin is possibly causing biotic homogenization. Thermophilic species, i.e. Indo-Pacific, Indian Ocean, Red Sea, Tropical Atlantic, Tropical Pacific, and circum(sub)tropical, account for 88.4% of the introduced species in the EMED, 72.8% in the CMED, 59.3% in the WMED and 56.1% in the Adriatic. Cold water species, i.e. circumboreal, N Atlantic, and N Pacific, make up a small percentage of the introduced species, ranging between 4.2% and 21.6% and being more numerous in the Adriatic and less so in the EMED. Species that are classified as invasive or potentially invasive are 134 in the whole of the Mediterranean: 108 are present in the EMED, 75 in the CMED, 53 in the Adriatic and 64 in the WMED. The WMED hosts most invasive macrophytes, whereas the EMED has the lion’s share in polychaetes, crustaceans, molluscs and fish.
RESUMEN: El estudio de los fondos de concrecionamiento coraligeno presenta un gran interes, no so... more RESUMEN: El estudio de los fondos de concrecionamiento coraligeno presenta un gran interes, no solo por la gran riqueza especifica existente sino tambien por representar un refugio en donde desarrollan toda o parte de su vida muchas especies. A pesar de esto, no existen estudios sobre Decapodos en los que de forma cuantitativa se aborde la es t ructura de la comunidad durante un ciclo anual. Por ello, el presente trabajo representa una pr imera contribucion al conocimiento de la comunidad, de este grupo, existente en los fondos superficiales (—3, —4 m) del alga calcarea Mesophyllum lichenoides (ELLIS & SOL.) Lemoine, de Malaga. De las 20 especies encontradas solo 6 pueden considerarse como las mas «caracteristicas» del biotopo estudiado. Estas son: Pilumnus hirtellus, Alpheus dentipes, Pisidia longimana, Synalpheus hululensis, Athanas nitescens y Cestopagurus timidus. Todas ellas presentan unos valores altos de constancia y abundancia durante el ano, aunque ninguna es exclusiva de e...
Bulletin of Marine Science, 1996
During the Balgim-84 Expedition, carried out from both sides of the Straits of Gibraltar (Atlanti... more During the Balgim-84 Expedition, carried out from both sides of the Straits of Gibraltar (Atlantic Ocean: Gulf of Cadiz and Mediterranean Sea: Alboran Sea), in depths between 137 and 2,142 m, 3,559 specimens of decapods belonging to 122 species were captured. Many of these species are rare, of biogeographical interest and represent new records for the area. For some species, anatomical details are given. A specimen of Plesionika may be a new species. The collection of Calocarides coronatus, Sympagurus acinops, Monodaeus guinotae, Periclimenes kornii, Alpheus talismani, Athanas amazone, Merhippolyte ancitrosta, Eualus cf. lebourae, Plesionika williamsi, Pontophilus cf. abyssi, Spongicoloides evolutus, Spongicoloides profundus, Odontozona edwardsi, Catapaguroides iris and Strobopagurus gracilipes, among others, represents new records for the Mediterranean Sea, eastern Atlantic, and Iberian waters.
Invertebrate Biology, 2005
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Cahiers De Biologie Marine, 1992
The collect of a male specimen of Micl'ocassiope lIIinol' (Dana) from the southeast of Spain repr... more The collect of a male specimen of Micl'ocassiope lIIinol' (Dana) from the southeast of Spain represe nts the first record of the species for the West Mediterranean Sea. The recent studies of the decapod faLma of the Alboran Sea show that West African Atl antic species are co lonizing thi s Mediterranean area. , Résumé: Le déve loppement récent des recherches sur les Décapodes dans la Mer de Alboran (à l' est de Gibraltar) montre l' extensio n d' espèces ouest-africains en Méditerranée. Microcassiope minol' (Dana) , récoltée dans des cavités d'algues corallinacées, est signalée pour la première fois sur les côtes européennes de la Méditerranée occidentale (Espagne).
Cahiers De Biologie Marine, 1992
The study of the evolution of a Crustacea Decapoda community from calcareous bottoms - strongly r... more The study of the evolution of a Crustacea Decapoda community from calcareous bottoms - strongly related with Posidonia oceanica, developed during 1986-1987. completes the overview of the structure and varia- bility existing in this habitat. Four groups have been stablished, according to the abundance-presence. The cualita- tive composition is similar to that found in other calcareous bottoms from the Southern Spain. So, the six more abundant species (belonging to the first and second groups) are the same. The fluctuations of the general structure and its evolution are controled by the two dominant species, Athanas nitescens and Pisidia longicomis (form. lon- gimana). The development of the Posidonia bed. which increases the sedimentation and protection, is, possibly, the responsible of the cuantitative differences found.
Scientia Marina
The unique colour patterns of the hermit crab species inhabiting the Iberian Peninsula and geogra... more The unique colour patterns of the hermit crab species inhabiting the Iberian Peninsula and geographically close areas are studied based on colour patterns observed in live specimens either in the field or live in the laboratory. Live colour patterns are shown to be useful for differentiating species in the Paguroidea from Iberian waters. Colour information has not been frequently documented in previous studies, mainly because of difficulties in accessing live specimens. Up to 51 species are currently recorded within the study area, 45 of which are included in the present work, focusing on the distinctive colour traits for each species that can be observed in the field. A complete key for identifying the species within the study area based on colour patterns is included. This study is the first attempt to put in place this type of tool oriented to field work. Further studies will be required to complete and update this information, especially for species which are scarce or difficult...
Scientia Marina, 2008
The structure of decapod crustacean assemblages living in shallow waters (5 to 25 m depth) in an ... more The structure of decapod crustacean assemblages living in shallow waters (5 to 25 m depth) in an area of the Alborán Sea (Mediterranean Sea) neighbouring the Strait of Gibraltar was studied. The relatively high richness found in this area is probably due to the diversity of substrata and the confluence of Atlantic and Mediterranean waters. The most abundant species was the hermit crab Diogenes pugilator due to the prevalence of sandy substrate, the shallow depth-range, and the species' life history. The coralligenous bottom showed the highest species richness and diversity values due to the complexity of microhabitats in this type of bottom and probably due to the food flux associated with currents. Conversely, shallow, well calibrated, fine to medium sandy bottoms of 5 m had the lowest values and the maximum densities due to the high abundance of a few well adapted species. The analysis of the different stations showed significant spatial differences according to depth and sedimentary characteristics. Depth is the environmental variable that correlates best with the decapod assemblages, with a particularly significant boundary between 5 m and 15 m. Nevertheless, there was a continuous transition between the assemblages. These results evidence the importance of quantitative studies in differentiating decapod assemblages. Relationships between these assemblages should also be taken into account in coastal management, since altering a substrate could have repercussions for the structure of the communities of neighbouring substrates. No seasonal significant differences were found in the overall analysis of the area, but there were differences between spring and autumn and spring and summer in relation to depth and substrate (crossed analysis). Finally, we present the species which allow us to discriminate the different assemblages according to sediment and depth, as well as the species' contributions.
Scientia Marina, 2002
The present paper presents an overview of the studies on the genus Paguristes from the Caribbean ... more The present paper presents an overview of the studies on the genus Paguristes from the Caribbean Sea and more specifically from Cuban waters. Additionally, the hermit crabs belonging to the genus Paguristes collected during different surveys of the "Instituto de Oceanología de Cuba" on the Cuban coast have been listed. A new species of hermit crab, Paguristes hologuinensis sp. nov., is described and remarks on a new record, Paguristes paraguanensis MacLaughlin and Provenzano, 1974, are given.
nean. Arxius de Miscel·lànea Zoològica, vol. 7: 72–77. New records of Parasquilla ferussaci (Roux... more nean. Arxius de Miscel·lànea Zoològica, vol. 7: 72–77. New records of Parasquilla ferussaci (Roux, 1830) (Crustacea, Stomatopoda) from the Eastern Atlantic and Western Mediterranean. — We report the occurrence of the little known stomatopod Parasquilla ferussaci on the Atlantic and Mediterranean coasts of the Iberian peninsula. Documentation is based on three specimens captured off Isla Cristina (Huelva) in the Gulf of Cadiz, off Fuengirola (Málaga) in the Alboran Sea and off Gavà (Barcelona) in the North–Western Mediterranean. These reports fill the distribution gap between Eastern Central Atlantic reports and previous Mediterranean reports east of the Balearic Islands.
Marine Protected Areas (MPAs) and Key Biodiversity Areas (KBAs) are important tools for preservin... more Marine Protected Areas (MPAs) and Key Biodiversity Areas (KBAs) are important tools for preserving species, habitats and ecological-geological-oceanographical processes. Nowadays, more than 10% of the Alboran Sea is under some protected figure or is on its way to be protected. This chapter presents the current state of the MPAs and KBAs of the Alboran Sea. Most of the habitats included in the Annex I of the Habitats Directive or the Barcelona Convention are represented in MPAs-KBAs of the Alboran Sea. More than 290 threatened species are also occurring in these MPAs-KBAs, but some of them have experienced a strong decline even within them (e.g. Zostera marina, Pinna nobilis, etc.). Although important progress in the protection of the Alboran Sea has been made in the last decades, there is still much to be done for effective conservation of the habitats and species. The main efforts may focus on improving (1) knowledge of the habitats and species, (2) the surveillance and mitigation against current impacts, (3) the coordination between local, national and international agencies and (4) the measures for improving habitats and threatened species populations, among others. A deep change in the socioeconomic model is then needed for preserving the Alboran Sea natural heritage, promoting then a sustainable use of its resources.
Mediterranean Marine Science, 2012
More than 60 marine non-indigenous species (NIS) have been removed from previous lists and 84 spe... more More than 60 marine non-indigenous species (NIS) have been removed from previous lists and 84 species have been added, bringing the total to 986 alien species in the Mediterranean [775 in the eastern Mediterranean (EMED), 249 in the central Mediterranean (CMED), 190 in the Adriatic Sea (ADRIA) and 308 in the western Mediterranean (WMED)]. There were 48 new entries since 2011 which can be interpreted as approximately one new entry every two weeks. The number of alien species continues to increase, by 2-3 species per year for macrophytes, molluscs and polychaetes, 3-4 species per year for crustaceans, and 6 species per year for fish. The dominant group among alien species is molluscs (with 215 species), followed by crustaceans (159) and polychaetes (132). Macrophytes are the leading group of NIS in the ADRIA and the WMED, reaching 26-30% of all aliens, whereas in the EMED they barely constitute 10% of the introductions. In the EMED, molluscs are the most species-rich group, followed b...
Journal of Crustacean Biology
Integrative taxonomy studies have allowed us to clarify some taxonomic problems in cryptic specie... more Integrative taxonomy studies have allowed us to clarify some taxonomic problems in cryptic species within species of InachusWeber, 1795 with two protogastric tubercles found in European waters. Several morphotypes of Inachus phalangium (Fabricius, 1775) are recognized, and a new species is described from the Atlantic coast of the Iberian Peninsula, for which distribution data are provided. Furthermore, two Inachus cf. thoracicus specimens with a unique sternal morphology were collected from Málaga Spain, but their preservation in formaldehyde prevented molecular analyses. Specimens of I. guentheri (Miers, 1879) have been re-examined and the validity of previous reports from tropical and subtropical Atlantic waters is discussed. These results allow us to separate and clarify the status of species within this group. An identification key for Inachus species with two protogastric tubercles is also provided.
Información del artículo Aportaciones al conocimiento de la fauna y flora litoral de la isla de A... more Información del artículo Aportaciones al conocimiento de la fauna y flora litoral de la isla de Alborán (España).
ABSTRACT The state-of-art on alien species in the Mediterranean Sea is presented, making distinct... more ABSTRACT The state-of-art on alien species in the Mediterranean Sea is presented, making distinctions among the four subregions defined in the EU Marine Strategy Framework Directive: (i) the Western Mediterranean Sea (WMED); (ii) the Central Mediterranean Sea (CMED); (iii) the Adriatic Sea (ADRIA); and (iv) the Eastern Mediterranean Sea (EMED). The updated checklist (December 2010) of marine alien species within each subregion, along with their acclimatization status and origin, is provided. A total of 955 alien species is known in the Mediterranean, the vast majority of them having being introduced in the EMED (718), less in the WMED (328) and CMED (267) and least in the Adriatic (171). Of these, 535 species (56%) are established in at least one area. Despite the collective effort of experts who attempted in this work, the number of introduced species remains probably underestimated. Excluding microalgae, for which knowledge is still insufficient, aliens have increased the total species richness of the Mediterranean Sea by 5.9%. This figure should not be directly read as an indication of higher biodiversity, as spreading of so many aliens within the basin is possibly causing biotic homogenization. Thermophilic species, i.e. Indo-Pacific, Indian Ocean, Red Sea, Tropical Atlantic, Tropical Pacific, and circum(sub)tropical, account for 88.4% of the introduced species in the EMED, 72.8% in the CMED, 59.3% in the WMED and 56.1% in the Adriatic. Cold water species, i.e. circumboreal, N Atlantic, and N Pacific, make up a small percentage of the introduced species, ranging between 4.2% and 21.6% and being more numerous in the Adriatic and less so in the EMED. Species that are classified as invasive or potentially invasive are 134 in the whole of the Mediterranean: 108 are present in the EMED, 75 in the CMED, 53 in the Adriatic and 64 in the WMED. The WMED hosts most invasive macrophytes, whereas the EMED has the lion’s share in polychaetes, crustaceans, molluscs and fish.
RESUMEN: El estudio de los fondos de concrecionamiento coraligeno presenta un gran interes, no so... more RESUMEN: El estudio de los fondos de concrecionamiento coraligeno presenta un gran interes, no solo por la gran riqueza especifica existente sino tambien por representar un refugio en donde desarrollan toda o parte de su vida muchas especies. A pesar de esto, no existen estudios sobre Decapodos en los que de forma cuantitativa se aborde la es t ructura de la comunidad durante un ciclo anual. Por ello, el presente trabajo representa una pr imera contribucion al conocimiento de la comunidad, de este grupo, existente en los fondos superficiales (—3, —4 m) del alga calcarea Mesophyllum lichenoides (ELLIS & SOL.) Lemoine, de Malaga. De las 20 especies encontradas solo 6 pueden considerarse como las mas «caracteristicas» del biotopo estudiado. Estas son: Pilumnus hirtellus, Alpheus dentipes, Pisidia longimana, Synalpheus hululensis, Athanas nitescens y Cestopagurus timidus. Todas ellas presentan unos valores altos de constancia y abundancia durante el ano, aunque ninguna es exclusiva de e...