Jorge Calvo | Universidad Nacional del Comahue (original) (raw)

Papers by Jorge Calvo

Research paper thumbnail of Rio de Janeiro, Brasil

A new Cretaceous terrestrial ecosystem from Gondwana with the description of a new sauropod dinosaur

Research paper thumbnail of An articulated titanosaur from Patagonia (Argentina): New evidence of neosauropod pedal evolution

Research paper thumbnail of A new Abelisauroid from the Upper Cretaceous of Brazil

Although important dinosaur specimens have been recently described from Brazil, the theropod reco... more Although important dinosaur specimens have been recently described from Brazil, the theropod record in this county is still rather scarce, particularly from Cretaceous strata. Here we describe a complete right tibia (MCT 1783-R) from the Marília Formation (Maastrichtian, Bauru Basin) near Peirópolis (Uberaba, Minas Gerais). This is the best theropod specimen recovered from this unit after five decades of collecting. MCT 1783-R shows a well developed cnemial crest with a lobular distal end, that projects significantly above the proximal articular surface, features that are characteristic of the Abelisauroidea. It further shows unique characters such as a marked asymmetry of lateral and medial condyles in posterior view, and a very deep triangular articular face for the astragalus, suggesting that it represents a new taxon.

Research paper thumbnail of Figure 1 in Anatomy of the basal titanosaur (Dinosauria, Sauropoda) Andesaurus delgadoi from the mid-Cretaceous (Albian-early Cenomanian) Río Limay Formation, Neuquén Province, Argentina: implications for titanosaur systematics

Figure 1. Map showing the El Chocón locality where the holotype of Andesaurus delgadoi was discov... more Figure 1. Map showing the El Chocón locality where the holotype of Andesaurus delgadoi was discovered (marked with a star). Inset map A shows the position of Neuquén Province within Argentina, and map B shows Neuquén Province. The black box within map B represents the area displayed in the main map (figures modified from Calvo, 1999).

Research paper thumbnail of Figure 7 in Anatomy of the basal titanosaur (Dinosauria, Sauropoda) Andesaurus delgadoi from the mid-Cretaceous (Albian-early Cenomanian) Río Limay Formation, Neuquén Province, Argentina: implications for titanosaur systematics

Figure 7. Andesaurus delgadoi. Photographs of middle-posterior caudal vertebrae: (A) Cd23 in left... more Figure 7. Andesaurus delgadoi. Photographs of middle-posterior caudal vertebrae: (A) Cd23 in left lateral view; (B) Cd24 in left lateral view; Cd25 in (C) anterior, (D) right lateral, and (E) posterior view. Scale bar: 100 mm.

Research paper thumbnail of Figure 4 in Anatomy of the basal titanosaur (Dinosauria, Sauropoda) Andesaurus delgadoi from the mid-Cretaceous (Albian-early Cenomanian) Río Limay Formation, Neuquén Province, Argentina: implications for titanosaur systematics

Figure 4. Andesaurus delgadoi. Photographs of anterior caudal vertebrae: (A) Cd2 in right lateral... more Figure 4. Andesaurus delgadoi. Photographs of anterior caudal vertebrae: (A) Cd2 in right lateral view; Cd3 in (B) left lateral, (C) dorsal, (D) anterior, (E) posterior, and (F) ventral views; vf, vascular foramen. Scale bar: 100 mm.

Research paper thumbnail of Figure 8 in Anatomy of the basal titanosaur (Dinosauria, Sauropoda) Andesaurus delgadoi from the mid-Cretaceous (Albian-early Cenomanian) Río Limay Formation, Neuquén Province, Argentina: implications for titanosaur systematics

Figure 8. Andesaurus delgadoi. Photographs of chevrons: (A) chevron 1 (C1) in anterior and right ... more Figure 8. Andesaurus delgadoi. Photographs of chevrons: (A) chevron 1 (C1) in anterior and right lateral view; (B) chevron 2 (C2) in anterior and left lateral view; (C) chevron 3 (C3) in anterior and left lateral view; (D) chevron 4 (C4) in posterior and left lateral view. Scale bar: 200 mm.

Research paper thumbnail of Figure 3 in Anatomy of the basal titanosaur (Dinosauria, Sauropoda) Andesaurus delgadoi from the mid-Cretaceous (Albian-early Cenomanian) Río Limay Formation, Neuquén Province, Argentina: implications for titanosaur systematics

Figure 3. Andesaurus delgadoi. Photographs of first caudal vertebra (Cd1) in (A) anterior, (B) po... more Figure 3. Andesaurus delgadoi. Photographs of first caudal vertebra (Cd1) in (A) anterior, (B) posterior, (C) left lateral, and (D) dorsal (anterior surface at top of image) view. Scale bar: 100 mm.

Research paper thumbnail of New material of Peirosaurids from Neuquén, Patagonia: its age

The first evidence of a crocodyform on the Futalognko quarry (Neuquén, Argentina) is presented. T... more The first evidence of a crocodyform on the Futalognko quarry (Neuquén, Argentina) is presented. The Portezuelo Formation has given in this site an extraordinary fossil record. The more impressive materials correspond to sauropod, theropod and ornithopod dinosaurs. However; it is also important the evidence of turtles, pterosaurus, crocodiles, fish and plants. In this paper we describe a crocodilyform tooth that is assigned to the Peirosauridae family, probably to Lomasuchus palpebrosus. The tooth comes from the Portezuelo Formation. Two more crocodilyforms are known from the area: Lomasuchus palpebrosus herein referred to the Portezuelo Formation (upper Turonian-lower Coniacian) and Peirosaurus tormini assigned to the Bajo de la Carpa Formation (upper Coniacian). ABSTRACT-NUEVO MATERIAL DE PEIROSAURIDO DE NEUQUÉN, PATAGONIA: SU EDAD. Presentamos la primer evidencia de un crocodylifome del yacimiento Futalognko (Neuquén, Argentina). La Formación Portezuelo ha dado un extraordinario r...

Research paper thumbnail of Gonzalez_Riga_Supplementary_Fig_S4_articulated_pes_cast.pdf

Research paper thumbnail of Supplementary Fig. S4 for González Riga et al. (2016), "A gigantic new dinosaur from Argentina and the evolution of the sauropod hind foot

Research paper thumbnail of FROM THE NEUQUÉN GROUP (LATE CRETACEOUS), PATAGONIA, ARGENTINA 1 (With 20 figures)

ABSTRACT: Titanosaurs are among the largest dinosaurs known to date. Here we describe the anatomy... more ABSTRACT: Titanosaurs are among the largest dinosaurs known to date. Here we describe the anatomy of Futalognkosaurus dukei, the most complete giant sauropod ever found. It comes from outcrops of the Portezuelo Formation at the Barreales lake, some 90 km northwest of Neuquén city (Patagonia). The specimen consists of a complete neck, dorsal vertebrae with ribs, pelvis, and one caudal vertebra. Futalognkosaurus dukei is a member of the Titanosauridae and belongs to the Lognkosauria, a clade that includes Mendozasaurus neguyelap and probably also the giant Puertasaurus reuili.

Research paper thumbnail of Theropod Dinosaur Trackways from the Lower Cretaceous of the

We describe sixteen theropod dinosaur trackways from site III (Lower Cretaceous?) of the Quebrada... more We describe sixteen theropod dinosaur trackways from site III (Lower Cretaceous?) of the Quebrada Chacarilla in the Chacarilla Formation (Jurassic-Lower Cretaceous), northern Chile. The trackbed belongs to a meandering river environment, with recurrent track assemblages. We fi nd that the main direction of the ichnites from site III parallels that of water fl ow. Two theropod footprint morphotypes are recognized: A) digit II and IV occupying about 0.5 times the total footprint length, and B) digit II and IV occupying less than 0.35 times the total footprint length. We estimate similar speed (~7 km/h) for the two parallel theropod trackways of morphotype A, suggesting synchro nized walking. We also describe some of the largest theropod footprints from South America (>60 cm), which are all of morphotype B.

Research paper thumbnail of Juvenile titanosaurian (Dinosauria: Sauropoda) remains from the Upper Cretaceous of Barreales Lake, Neuquén, Argentina

Research paper thumbnail of An overview of Upper Cretaceous dinosaur tracks and other trace fossils from the Barreales lake area, Neuquén Province, Patagonia, Argentina

Journal of South American Earth Sciences

Research paper thumbnail of Cranial Osteology of Amargasaurus Cazaui Salgado & Bonaparte (Sauropoda, Dicraeosauridae) from the Neocomian of Patagonia

Ameghiniana, 2013

Se describe la anatomia craneana del Dicraeosaurido Amargasaurus cazauiui. El craneo de Amargasau... more Se describe la anatomia craneana del Dicraeosaurido Amargasaurus cazauiui. El craneo de Amargasaurus cazaui es semejante al de Dicraesaurus hansemanni. Se confirma la presencia de fontanelas parietales y postparietales en el techo craneano de los dicraesarauridos. La evidencia muestra que los dicraeosauridos tenian un craneo mas derivado que el de los Apatosaurus, Nemegtosaurus y Quaesitosaurus se excluyen de los Dicraeosauridae porque ellos no comparten probables rasgos derivados con Dicraeosaurus y Amargasaurus.

Research paper thumbnail of Análisis tafonómicos de saurópodos titanosaurios en la Cuenca Neuquina (Argentina) y sus implicaciones en estudios sistemáticos

Research paper thumbnail of A new Abelisauridae (Dinosauria: Theropoda) from northwest Patagonia

Abstract. A new theropod Abelisauridae is described, Ekrixinatosaurusnovasi gen. et sp. nov., of ... more Abstract. A new theropod Abelisauridae is described, Ekrixinatosaurusnovasi gen. et sp. nov., of northwestern Patagonia (Neuquen Province, Argentina). A preliminar cladistic analysis placed Ekrixinatosaurus together with Majungatholus and Carnotaurini; and the sister group is Ilokelesia. The presence of this specimen in Albian-Cenomanian beds allows us to support the early divergence (pre-Senonian) of this clade, and the hypothesis of the presence of Abelisauridae in continental Africa. The present distribution of Abelisauridae indicates: 1) a vicariance of this clade based on a pre-Cenomanian pan-Gondwanic distribution; 2) dispersion throughout terrestrial bridges. In both cases, the absence of Abelisauridae in continental Africa can be considered a bias of the fossil record. Resumen. UN NUEVO ABELISAURIDAE (DINOSAURIA: THEROPODA) DEL NOROESTE DE LA PATAGONIA. Se describe un nuevo teropodo Abelisauridae, Ekrixinatosaurusnovasi gen. et sp. nov., del noroeste de la Patagonia (provinc...

Research paper thumbnail of Huellas de dinosaurios en la costa oeste del embalse Ezequiel Ramos Mexía y alrededores (Cretácico Superior, Provincia de Neuquén, República Argentina)

Boletín de la Sociedad Geológica Mexicana

In Argentina, some of the most important dinosaur tracksites are found on the west coast of the E... more In Argentina, some of the most important dinosaur tracksites are found on the west coast of the Ezequiel Ramos Mexía reservoir, in the Neuquén province. In the last 30 years, more than 100 km of coastline have yielded hundreds of dinosaur tracks. The tracks are part of the Albian?-lower Cenomanian Candeleros Formation, Neuquén Group. An extensive study and review of the dinosaur tracks described in the past 30 years is reinterpreted and revised. Candeleroichnus canalei ichnogen. nov. ichnosp. nov. is described. A neotype is chosen for Deferrariischnium mapuchensis Calvo, 1991 and an emended diagnosis improves the ichnospecies. In addition, new information of the trackway diagnosis of Picunichnus benedettoi Calvo, 1991, such as length of pace and pace angle is given. We analyze all the dinosaur sites discovered until now and we identify all described ichnoforms such as Sousaichnium monettae, Limayichnus major, Abelichnus astigarrae and Sauropodichnus giganteus. We include unpublished figures, photos and tables of the ichnotypes.

Research paper thumbnail of Evolution of Titanosaurid Sauropods. II: The Cranial Evidence

Ameghiniana, Feb 6, 2014

... II: THE CRANIAL EVIDENCE Leonardo SALGADO'and Jorge Orlando CALVO' ABSTRACT... more ... II: THE CRANIAL EVIDENCE Leonardo SALGADO'and Jorge Orlando CALVO' ABSTRACT. ... Many traits of the Diplodocus-lype skull are correlat-ed. For instance, in Diplodocus (see Berman andMcIntosh, l978, fig. 9) and Camarasaurus Cope (see Osborn and Mook, l92l, pl. ...

Research paper thumbnail of Rio de Janeiro, Brasil

A new Cretaceous terrestrial ecosystem from Gondwana with the description of a new sauropod dinosaur

Research paper thumbnail of An articulated titanosaur from Patagonia (Argentina): New evidence of neosauropod pedal evolution

Research paper thumbnail of A new Abelisauroid from the Upper Cretaceous of Brazil

Although important dinosaur specimens have been recently described from Brazil, the theropod reco... more Although important dinosaur specimens have been recently described from Brazil, the theropod record in this county is still rather scarce, particularly from Cretaceous strata. Here we describe a complete right tibia (MCT 1783-R) from the Marília Formation (Maastrichtian, Bauru Basin) near Peirópolis (Uberaba, Minas Gerais). This is the best theropod specimen recovered from this unit after five decades of collecting. MCT 1783-R shows a well developed cnemial crest with a lobular distal end, that projects significantly above the proximal articular surface, features that are characteristic of the Abelisauroidea. It further shows unique characters such as a marked asymmetry of lateral and medial condyles in posterior view, and a very deep triangular articular face for the astragalus, suggesting that it represents a new taxon.

Research paper thumbnail of Figure 1 in Anatomy of the basal titanosaur (Dinosauria, Sauropoda) Andesaurus delgadoi from the mid-Cretaceous (Albian-early Cenomanian) Río Limay Formation, Neuquén Province, Argentina: implications for titanosaur systematics

Figure 1. Map showing the El Chocón locality where the holotype of Andesaurus delgadoi was discov... more Figure 1. Map showing the El Chocón locality where the holotype of Andesaurus delgadoi was discovered (marked with a star). Inset map A shows the position of Neuquén Province within Argentina, and map B shows Neuquén Province. The black box within map B represents the area displayed in the main map (figures modified from Calvo, 1999).

Research paper thumbnail of Figure 7 in Anatomy of the basal titanosaur (Dinosauria, Sauropoda) Andesaurus delgadoi from the mid-Cretaceous (Albian-early Cenomanian) Río Limay Formation, Neuquén Province, Argentina: implications for titanosaur systematics

Figure 7. Andesaurus delgadoi. Photographs of middle-posterior caudal vertebrae: (A) Cd23 in left... more Figure 7. Andesaurus delgadoi. Photographs of middle-posterior caudal vertebrae: (A) Cd23 in left lateral view; (B) Cd24 in left lateral view; Cd25 in (C) anterior, (D) right lateral, and (E) posterior view. Scale bar: 100 mm.

Research paper thumbnail of Figure 4 in Anatomy of the basal titanosaur (Dinosauria, Sauropoda) Andesaurus delgadoi from the mid-Cretaceous (Albian-early Cenomanian) Río Limay Formation, Neuquén Province, Argentina: implications for titanosaur systematics

Figure 4. Andesaurus delgadoi. Photographs of anterior caudal vertebrae: (A) Cd2 in right lateral... more Figure 4. Andesaurus delgadoi. Photographs of anterior caudal vertebrae: (A) Cd2 in right lateral view; Cd3 in (B) left lateral, (C) dorsal, (D) anterior, (E) posterior, and (F) ventral views; vf, vascular foramen. Scale bar: 100 mm.

Research paper thumbnail of Figure 8 in Anatomy of the basal titanosaur (Dinosauria, Sauropoda) Andesaurus delgadoi from the mid-Cretaceous (Albian-early Cenomanian) Río Limay Formation, Neuquén Province, Argentina: implications for titanosaur systematics

Figure 8. Andesaurus delgadoi. Photographs of chevrons: (A) chevron 1 (C1) in anterior and right ... more Figure 8. Andesaurus delgadoi. Photographs of chevrons: (A) chevron 1 (C1) in anterior and right lateral view; (B) chevron 2 (C2) in anterior and left lateral view; (C) chevron 3 (C3) in anterior and left lateral view; (D) chevron 4 (C4) in posterior and left lateral view. Scale bar: 200 mm.

Research paper thumbnail of Figure 3 in Anatomy of the basal titanosaur (Dinosauria, Sauropoda) Andesaurus delgadoi from the mid-Cretaceous (Albian-early Cenomanian) Río Limay Formation, Neuquén Province, Argentina: implications for titanosaur systematics

Figure 3. Andesaurus delgadoi. Photographs of first caudal vertebra (Cd1) in (A) anterior, (B) po... more Figure 3. Andesaurus delgadoi. Photographs of first caudal vertebra (Cd1) in (A) anterior, (B) posterior, (C) left lateral, and (D) dorsal (anterior surface at top of image) view. Scale bar: 100 mm.

Research paper thumbnail of New material of Peirosaurids from Neuquén, Patagonia: its age

The first evidence of a crocodyform on the Futalognko quarry (Neuquén, Argentina) is presented. T... more The first evidence of a crocodyform on the Futalognko quarry (Neuquén, Argentina) is presented. The Portezuelo Formation has given in this site an extraordinary fossil record. The more impressive materials correspond to sauropod, theropod and ornithopod dinosaurs. However; it is also important the evidence of turtles, pterosaurus, crocodiles, fish and plants. In this paper we describe a crocodilyform tooth that is assigned to the Peirosauridae family, probably to Lomasuchus palpebrosus. The tooth comes from the Portezuelo Formation. Two more crocodilyforms are known from the area: Lomasuchus palpebrosus herein referred to the Portezuelo Formation (upper Turonian-lower Coniacian) and Peirosaurus tormini assigned to the Bajo de la Carpa Formation (upper Coniacian). ABSTRACT-NUEVO MATERIAL DE PEIROSAURIDO DE NEUQUÉN, PATAGONIA: SU EDAD. Presentamos la primer evidencia de un crocodylifome del yacimiento Futalognko (Neuquén, Argentina). La Formación Portezuelo ha dado un extraordinario r...

Research paper thumbnail of Gonzalez_Riga_Supplementary_Fig_S4_articulated_pes_cast.pdf

Research paper thumbnail of Supplementary Fig. S4 for González Riga et al. (2016), "A gigantic new dinosaur from Argentina and the evolution of the sauropod hind foot

Research paper thumbnail of FROM THE NEUQUÉN GROUP (LATE CRETACEOUS), PATAGONIA, ARGENTINA 1 (With 20 figures)

ABSTRACT: Titanosaurs are among the largest dinosaurs known to date. Here we describe the anatomy... more ABSTRACT: Titanosaurs are among the largest dinosaurs known to date. Here we describe the anatomy of Futalognkosaurus dukei, the most complete giant sauropod ever found. It comes from outcrops of the Portezuelo Formation at the Barreales lake, some 90 km northwest of Neuquén city (Patagonia). The specimen consists of a complete neck, dorsal vertebrae with ribs, pelvis, and one caudal vertebra. Futalognkosaurus dukei is a member of the Titanosauridae and belongs to the Lognkosauria, a clade that includes Mendozasaurus neguyelap and probably also the giant Puertasaurus reuili.

Research paper thumbnail of Theropod Dinosaur Trackways from the Lower Cretaceous of the

We describe sixteen theropod dinosaur trackways from site III (Lower Cretaceous?) of the Quebrada... more We describe sixteen theropod dinosaur trackways from site III (Lower Cretaceous?) of the Quebrada Chacarilla in the Chacarilla Formation (Jurassic-Lower Cretaceous), northern Chile. The trackbed belongs to a meandering river environment, with recurrent track assemblages. We fi nd that the main direction of the ichnites from site III parallels that of water fl ow. Two theropod footprint morphotypes are recognized: A) digit II and IV occupying about 0.5 times the total footprint length, and B) digit II and IV occupying less than 0.35 times the total footprint length. We estimate similar speed (~7 km/h) for the two parallel theropod trackways of morphotype A, suggesting synchro nized walking. We also describe some of the largest theropod footprints from South America (>60 cm), which are all of morphotype B.

Research paper thumbnail of Juvenile titanosaurian (Dinosauria: Sauropoda) remains from the Upper Cretaceous of Barreales Lake, Neuquén, Argentina

Research paper thumbnail of An overview of Upper Cretaceous dinosaur tracks and other trace fossils from the Barreales lake area, Neuquén Province, Patagonia, Argentina

Journal of South American Earth Sciences

Research paper thumbnail of Cranial Osteology of Amargasaurus Cazaui Salgado & Bonaparte (Sauropoda, Dicraeosauridae) from the Neocomian of Patagonia

Ameghiniana, 2013

Se describe la anatomia craneana del Dicraeosaurido Amargasaurus cazauiui. El craneo de Amargasau... more Se describe la anatomia craneana del Dicraeosaurido Amargasaurus cazauiui. El craneo de Amargasaurus cazaui es semejante al de Dicraesaurus hansemanni. Se confirma la presencia de fontanelas parietales y postparietales en el techo craneano de los dicraesarauridos. La evidencia muestra que los dicraeosauridos tenian un craneo mas derivado que el de los Apatosaurus, Nemegtosaurus y Quaesitosaurus se excluyen de los Dicraeosauridae porque ellos no comparten probables rasgos derivados con Dicraeosaurus y Amargasaurus.

Research paper thumbnail of Análisis tafonómicos de saurópodos titanosaurios en la Cuenca Neuquina (Argentina) y sus implicaciones en estudios sistemáticos

Research paper thumbnail of A new Abelisauridae (Dinosauria: Theropoda) from northwest Patagonia

Abstract. A new theropod Abelisauridae is described, Ekrixinatosaurusnovasi gen. et sp. nov., of ... more Abstract. A new theropod Abelisauridae is described, Ekrixinatosaurusnovasi gen. et sp. nov., of northwestern Patagonia (Neuquen Province, Argentina). A preliminar cladistic analysis placed Ekrixinatosaurus together with Majungatholus and Carnotaurini; and the sister group is Ilokelesia. The presence of this specimen in Albian-Cenomanian beds allows us to support the early divergence (pre-Senonian) of this clade, and the hypothesis of the presence of Abelisauridae in continental Africa. The present distribution of Abelisauridae indicates: 1) a vicariance of this clade based on a pre-Cenomanian pan-Gondwanic distribution; 2) dispersion throughout terrestrial bridges. In both cases, the absence of Abelisauridae in continental Africa can be considered a bias of the fossil record. Resumen. UN NUEVO ABELISAURIDAE (DINOSAURIA: THEROPODA) DEL NOROESTE DE LA PATAGONIA. Se describe un nuevo teropodo Abelisauridae, Ekrixinatosaurusnovasi gen. et sp. nov., del noroeste de la Patagonia (provinc...

Research paper thumbnail of Huellas de dinosaurios en la costa oeste del embalse Ezequiel Ramos Mexía y alrededores (Cretácico Superior, Provincia de Neuquén, República Argentina)

Boletín de la Sociedad Geológica Mexicana

In Argentina, some of the most important dinosaur tracksites are found on the west coast of the E... more In Argentina, some of the most important dinosaur tracksites are found on the west coast of the Ezequiel Ramos Mexía reservoir, in the Neuquén province. In the last 30 years, more than 100 km of coastline have yielded hundreds of dinosaur tracks. The tracks are part of the Albian?-lower Cenomanian Candeleros Formation, Neuquén Group. An extensive study and review of the dinosaur tracks described in the past 30 years is reinterpreted and revised. Candeleroichnus canalei ichnogen. nov. ichnosp. nov. is described. A neotype is chosen for Deferrariischnium mapuchensis Calvo, 1991 and an emended diagnosis improves the ichnospecies. In addition, new information of the trackway diagnosis of Picunichnus benedettoi Calvo, 1991, such as length of pace and pace angle is given. We analyze all the dinosaur sites discovered until now and we identify all described ichnoforms such as Sousaichnium monettae, Limayichnus major, Abelichnus astigarrae and Sauropodichnus giganteus. We include unpublished figures, photos and tables of the ichnotypes.

Research paper thumbnail of Evolution of Titanosaurid Sauropods. II: The Cranial Evidence

Ameghiniana, Feb 6, 2014

... II: THE CRANIAL EVIDENCE Leonardo SALGADO'and Jorge Orlando CALVO' ABSTRACT... more ... II: THE CRANIAL EVIDENCE Leonardo SALGADO'and Jorge Orlando CALVO' ABSTRACT. ... Many traits of the Diplodocus-lype skull are correlat-ed. For instance, in Diplodocus (see Berman andMcIntosh, l978, fig. 9) and Camarasaurus Cope (see Osborn and Mook, l92l, pl. ...