Octávio Mateus | Universidade Nova de Lisboa (original) (raw)
Papers by Octávio Mateus
Although diplodocoid sauropods from Africa and the Americas are well known, their European record... more Although diplodocoid sauropods from Africa and the Americas are well known, their European record remains largely neglected. Here we redescribe Dinheirosaurus lourinhanensis from the Late Jurassic of Portugal. The holotype comprises two posterior cervical vertebrae, the dorsal series and a caudal centrum. Redescription demonstrates its validity on the basis of three autapomorphies: (1) posteriorly restricted ventral keel on posterior cervical vertebrae; (2) three small subcircular fossae posterior to the lateral coel on posterior cervical neural spines; (3) accessory lamina linking the hyposphene with base of the posterior centrodiapophyseal lamina in middle-posterior dorsal vertebrae. Phylogenetic analysis places Dinheirosaurus as the sister taxon to Supersaurus, and this clade forms the sister taxon to other diplodocines. However, this position should be treated with caution as Dinheirosaurus displays several plesiomorphic features absent in other diplodocids (including unbifurcated presacral neural spines, and dorsolaterally projecting diapophyses on dorsal vertebrae) and only four additional steps are required to place Dinheirosaurus outside of Flagellicaudata. We identify Amazonsaurus as the basal-most rebbachisaurid and recover Zapalasaurus outside of the South American Limaysaurinae, suggesting the biogeographic history of rebbachisaurids is more complex than previously proposed. Review of the European diplodocoid record reveals evidence for the earliest known diplodocid, as well as additional diplodocid remains from the Late Jurassic of Spain. A Portuguese specimen, previously referred to Dinheirosaurus, displays strong similarities to Apatosaurus from the contemporaneous Morrison Formation of North America, indicating the presence of a second Late Jurassic Portuguese diplodocid taxon. Along with Dinheirosaurus, these Portuguese remains provide further evidence for a Late Jurassic palaeobiogeographic connection between Europe and North America. No dicraeosaurids are currently known from Europe, but rebbachisaurids are present in the Early Cretaceous, with weak evidence for the earliest known representative from the Late Jurassic of Spain; however, more complete material is required to recognize early members of this clade.
Forty partial fossil skulls belonging to beaked whales (Cetacea, Odontoceti, Ziphiidae) were coll... more Forty partial fossil skulls belonging to beaked whales (Cetacea, Odontoceti, Ziphiidae) were collected by trawling and long-line fishing on Neogene (probably Late Early to Middle Miocene) layers of the Atlantic floor off the coasts of Portugal and Spain (Asturias and Galicia). The systematic study of the most diagnostic Iberian specimens, those preserving the rostrum and the dorsal part of the cranium, led to the recognition of two new genera (Globicetus n. gen. and Imocetus n. gen.) and four new species (Choneziphius leidyi n. sp., G. hiberus n. gen., n. sp., I. piscatus n. gen., n. sp., and Tusciziphius atlanticus n. sp.). Based on the matrix of a previous work, the phylogenetic analysis places all the new taxa in the subfamily Ziphiinae Gray, 1850. More fragmentary specimens are tentatively referred to the genera Caviziphius Bianucci & Post, 2005 and Ziphirostrum du Bus, 1868. Among these new ziphiids, extremely bizarre skull morphologies are observed. In G. hiberus n. gen., n. sp. the proximal portion of the rostrum bears a voluminous premaxillary spheroid. In T. atlanticus n. sp. a medial premaxillary bulge is present on the rostrum; together with asymmetric rostral maxillary eminences at the rostrum base, this bulge displays various degrees of elevation in different specimens, which may be interpreted as sexual dimorphism. Specimens of I. piscatus n. gen., n. sp. bear two sets of even crests: spur-like rostral maxillary crests and longitudinal maxillary crests laterally bordering a wide and long facial basin. A preliminary macroscopic observation of these elements indicates very dense bones, with a compactness comparable with that of cetacean ear bones. Questioning their function, the high medial rostral elements (the premaxillary spheroid of G. hiberus n. gen., n. sp. and the medial bulge of T. atlanticus n. sp.) remind the huge rostral maxillary crests of adult males of the extant Hyperoodon ampullatus (Forster, 1770). In the latter, the crests are very likely related to head-butting. However, they are made of much more spongy bone than in the fossil taxa studied here, and therefore possibly better mechanically suited for facing impacts. Other interpretations of these unusual bone specializations, related to deep-diving (ballast) and echolocation (sound reflection), fail to explain the diversity of shapes and the hypothetical sexual dimorphism observed in at least part of the taxa. The spur-like rostral maxillary crests and long maxillary crests limiting the large facial basin in I. piscatus n. gen., n. sp. and the excrescences on the maxilla at the rostrum base in Choneziphius spp. are instead interpreted as areas of origin for rostral and facial muscles, acting on the nasal passages, blowhole, and melon. From a palaeobiogeographic point of view, the newly described taxa further emphasize the differences in the North Atlantic (including Iberian Peninsula) and South African Neogene ziphiid faunal lists. Even if the stratigraphic context is poorly understood, leaving open the question of the geological age for most of the dredged specimens, these differences in the composition of cold to temperate northern and southern hemisphere fossil ziphiid faunas may be explained by a warm-water equatorial barrier.
Acta Palaeontologica Polonica, Dec 1, 2009
Netherlands Journal of Geosciences - Geologie en Mijnbouw, 2015
Stable oxygen isotope values of inoceramid marine bivalve shells recovered from Bentiaba, Angola,... more Stable oxygen isotope values of inoceramid marine bivalve shells recovered from Bentiaba, Angola, are utilised as a proxy for paleotemperatures during the Late Cretaceous development of the African margin of the South Atlantic Ocean. The d 18 O values derived from inoceramids show a long-term increase from -3.2‰ in the Late Turonian to values between -0.8 and -1.8‰ in the Late Campanian. Assuming a constant oceanic d 18 O value, an ∼2‰ increase may reflect cooling of the shallow marine environment at Bentiaba by approximately 10°. Bentiaba values are offset by about +1‰ from published records for bathyal Inoceramus at Walvis Ridge. This offset in d 18 O values suggests a temperature difference of ∼5°b etween coastal and deeper water offshore Angola. Cooler temperatures implied by the d 18 O curve at Bentiaba coincide with the stratigraphic distribution of diverse marine amniotes, including mosasaurs, at Bentiaba.
Netherlands Journal of Geosciences - Geologie en Mijnbouw, 2015
ٰۣۧۙۗۢۙۗۧۙڷۣۚڷ۠ٷۢۦ۩ۣЂڷۧۘۢٷ۠ۦۙۜۨۙІ ۜۨۨۤۃҖҖ۞ۣ۩ۧ۠ٷۢۦғۗۙۛۘۦۖۡٷғۣۛۦҖІЂٰ ڷۦۣۚڷ۪ۧۙۗۦۙۧڷ۠ٷ... more ٰۣۧۙۗۢۙۗۧۙڷۣۚڷ۠ٷۢۦ۩ۣЂڷۧۘۢٷ۠ۦۙۜۨۙІ ۜۨۨۤۃҖҖ۞ۣ۩ۧ۠ٷۢۦғۗۙۛۘۦۖۡٷғۣۛۦҖІЂٰ ڷۦۣۚڷ۪ۧۙۗۦۙۧڷ۠ٷۣۢۨۘۘۆẰếẳẰẽặẬẹắẾΝẺỀẽẹẬặΝẺằΝύẰẺẾẮẴẰẹẮẰẾ ۙۦۙۜڷ۟ۗ۠Өڷۃۧۨۦۙ۠ٷڷ۠ٷۡٮ ۙۦۙۜڷ۟ۗ۠Өڷۃۣۧۢۨۤۦۗۧۖ۩ۑ ۙۦۙۜڷ۟ۗ۠Өڷۃۧۨۢۦۤۙۦڷ۠ٷۗۦۣۙۡۡӨ ۙۦۙۜڷ۟ۗ۠Өڷۃڷۙۧ۩ڷۣۚڷۧۡۦۙے ۺ۠ۦٷٮڷۙۜۨڷۣۡۦۚڷۧۢۙۡۗۙۤۧڷۦ۩ٷۣۧۧۙ۠ۤڷۘۦ۩ٷۣۧۡۧٷ۠ۙڷۣۙۢۨۗۙۢۨۧۦٷڷ۫ۙІ ۧۦ۩ٷۣۧۧۙ۠ۤڷۢڷۡۧۜۤۦۣۣۡۘۙٷۤڷۣۢڷۣۧۨۢۙۡۡۗڷۘۢٷڷٷۣ۠ۛۢۆڷۣۚڷۢٷۨۜۗۦۨۧٷٷی ۧٷۦۣیڷғۋғҒیڷۘۢٷڷ۪ۧۙ۠ٷٰۣۢçڷۣۤۡ۠íۍڷғۆڷۃۧ۩ۙۨٷیڷғۍڷۃۤ۠۩ۜۗۑڷғۑғۆڷۃۣۧۖۗٷЂڷғۋғۋڷۃۢۙۦۛۘۢۋڷЂғڷۃۢۺۣۗ۠ێڷғЂғیڷۃۣ۞úٷۦۆڷғې ڿڽڷҒڷڽڷۤۤڷۃҢڽڼھڷۺۦٷ۩ۦۖۙٯڷҖڷۙ۠ۗۨۦۆڷόẴẽẾếạẴẰỂڷҖڷٰۣۧۙۗۢۙۗۧۙڷۣۚڷ۠ٷۢۦ۩ۣЂڷۧۘۢٷ۠ۦۙۜۨۙІ Ңڽڼھڷۺۦٷ۩ۦۖۙٯڷہڽڷۃۣۙۢ۠ۢڷۘۙۜۧ۠ۖ۩ێڷۃڿۀғۀڽڼھҖۢ۞ۛғۀڽڼڽғڼڽڷۃٲۍө ھڿۀڼڼڼۀڽڿۀۀۀڿڽڼڼۑٵۨۗٷۦۨۧۖٷۛҖۦۣۘۛۙғۦۖۡٷۗ۠ۧғٷۢۦ۩ۣ۞ҖҖۃۤۨۨۜڷۃۙ۠ۗۨۦٷڷۧۜۨڷۣۨڷ۟ۢۋ ۃۙ۠ۗۨۦٷڷۧۜۨڷۙۨۗڷۣۨڷۣ۫ٱ ۫ۙІڷۧٷۦۣیڷғۋғҒیڷۘۢٷڷ۪ۧۙ۠ٷٰۣۢçڷۣۤۡ۠íۍڷғۆڷۃۧ۩ۙۨٷیڷғۍڷۃۤ۠۩ۜۗۑڷғۑғۆڷۃۣۧۖۗٷЂڷғۋғۋڷۃۢۙۦۛۘۢۋڷЂғڷۃۢۺۣۗ۠ێڷғЂғیڷۃۣ۞úٷۦۆڷғې ۡۧۜۤۦۣۣۡۘۙٷۤڷۣۢڷۣۧۨۢۙۡۡۗڷۘۢٷڷٷۣ۠ۛۢۆڷۣۚڷۢٷۨۜۗۦۨۧٷٷیڷۺ۠ۦٷٮڷۙۜۨڷۣۡۦۚڷۧۢۙۡۗۙۤۧڷۦ۩ٷۣۧۧۙ۠ۤڷۘۦ۩ٷۣۧۡۧٷ۠ۙڷۣۙۢۨۗۙۢۨۧۦٷ ڿۀғۀڽڼھҖۢ۞ۛғۀڽڼڽғڼڽۃۣۘڷҢڽڼھڷۍӨЂڷۣۢڷۙ۠ۖٷ۠ٷ۪ۆڷۃٰۣۧۙۗۢۙۗۧۙڷۣۚڷ۠ٷۢۦ۩ۣЂڷۧۘۢٷ۠ۦۙۜۨۙІڷۧғۦ۩ٷۣۧۧۙ۠ۤڷۢ ۙۦۙۜڷ۟ۗ۠Өڷۃڷۣۧۢۧۧۡۦۙێڷۨۧۙ۩ۥۙې ҢڽڼھڷۖۙٯڷۂڽڷۣۢڷھڽғھڽھғڼڿڽғہہڷۃۧۧۙۦۘۘٷڷێٲڷۃٰۛҖІЂۦۣۘۛۙғۦۖۡٷۗ۠ۧғٷۢۦ۩ۣ۞ҖҖۃۤۨۨۜڷۣۡۦۚڷۘۙۘٷۣۣ۠ۢ۫ө Netherlands Journal of Geosciences --Geologie en Mijnbouw | P age 1 of 16.
ABSTRACT Eggshells from three extant crocodilian species – Crocodylus mindorensis, Paleosuchus pa... more ABSTRACT Eggshells from three extant crocodilian species – Crocodylus mindorensis, Paleosuchus palpebrosus, and Alligator mississippiensis – has been analyzed and described. A comparison between these and other extant and fossil crocodilian eggs showed that the egg external surface ornamentation may vary between the anastomo– and the ramotuberculate (sensu Carpenter 1999) types, and the newly described rugosocavate type, characterized by an irregularly rugose surface scattered by subcircular pits that not always correspond to pore openings (Marzola et al. 2014). [...]
Although diplodocoid sauropods from Africa and the Americas are well known, their European record... more Although diplodocoid sauropods from Africa and the Americas are well known, their European record remains largely neglected. Here we redescribe Dinheirosaurus lourinhanensis from the Late Jurassic of Portugal. The holotype comprises two posterior cervical vertebrae, the dorsal series and a caudal centrum. Redescription demonstrates its validity on the basis of three autapomorphies: (1) posteriorly restricted ventral keel on posterior cervical vertebrae; (2) three small subcircular fossae posterior to the lateral coel on posterior cervical neural spines; (3) accessory lamina linking the hyposphene with base of the posterior centrodiapophyseal lamina in middle-posterior dorsal vertebrae. Phylogenetic analysis places Dinheirosaurus as the sister taxon to Supersaurus, and this clade forms the sister taxon to other diplodocines. However, this position should be treated with caution as Dinheirosaurus displays several plesiomorphic features absent in other diplodocids (including unbifurcated presacral neural spines, and dorsolaterally projecting diapophyses on dorsal vertebrae) and only four additional steps are required to place Dinheirosaurus outside of Flagellicaudata. We identify Amazonsaurus as the basal-most rebbachisaurid and recover Zapalasaurus outside of the South American Limaysaurinae, suggesting the biogeographic history of rebbachisaurids is more complex than previously proposed. Review of the European diplodocoid record reveals evidence for the earliest known diplodocid, as well as additional diplodocid remains from the Late Jurassic of Spain. A Portuguese specimen, previously referred to Dinheirosaurus, displays strong similarities to Apatosaurus from the contemporaneous Morrison Formation of North America, indicating the presence of a second Late Jurassic Portuguese diplodocid taxon. Along with Dinheirosaurus, these Portuguese remains provide further evidence for a Late Jurassic palaeobiogeographic connection between Europe and North America. No dicraeosaurids are currently known from Europe, but rebbachisaurids are present in the Early Cretaceous, with weak evidence for the earliest known representative from the Late Jurassic of Spain; however, more complete material is required to recognize early members of this clade.
Forty partial fossil skulls belonging to beaked whales (Cetacea, Odontoceti, Ziphiidae) were coll... more Forty partial fossil skulls belonging to beaked whales (Cetacea, Odontoceti, Ziphiidae) were collected by trawling and long-line fishing on Neogene (probably Late Early to Middle Miocene) layers of the Atlantic floor off the coasts of Portugal and Spain (Asturias and Galicia). The systematic study of the most diagnostic Iberian specimens, those preserving the rostrum and the dorsal part of the cranium, led to the recognition of two new genera (Globicetus n. gen. and Imocetus n. gen.) and four new species (Choneziphius leidyi n. sp., G. hiberus n. gen., n. sp., I. piscatus n. gen., n. sp., and Tusciziphius atlanticus n. sp.). Based on the matrix of a previous work, the phylogenetic analysis places all the new taxa in the subfamily Ziphiinae Gray, 1850. More fragmentary specimens are tentatively referred to the genera Caviziphius Bianucci & Post, 2005 and Ziphirostrum du Bus, 1868. Among these new ziphiids, extremely bizarre skull morphologies are observed. In G. hiberus n. gen., n. sp. the proximal portion of the rostrum bears a voluminous premaxillary spheroid. In T. atlanticus n. sp. a medial premaxillary bulge is present on the rostrum; together with asymmetric rostral maxillary eminences at the rostrum base, this bulge displays various degrees of elevation in different specimens, which may be interpreted as sexual dimorphism. Specimens of I. piscatus n. gen., n. sp. bear two sets of even crests: spur-like rostral maxillary crests and longitudinal maxillary crests laterally bordering a wide and long facial basin. A preliminary macroscopic observation of these elements indicates very dense bones, with a compactness comparable with that of cetacean ear bones. Questioning their function, the high medial rostral elements (the premaxillary spheroid of G. hiberus n. gen., n. sp. and the medial bulge of T. atlanticus n. sp.) remind the huge rostral maxillary crests of adult males of the extant Hyperoodon ampullatus (Forster, 1770). In the latter, the crests are very likely related to head-butting. However, they are made of much more spongy bone than in the fossil taxa studied here, and therefore possibly better mechanically suited for facing impacts. Other interpretations of these unusual bone specializations, related to deep-diving (ballast) and echolocation (sound reflection), fail to explain the diversity of shapes and the hypothetical sexual dimorphism observed in at least part of the taxa. The spur-like rostral maxillary crests and long maxillary crests limiting the large facial basin in I. piscatus n. gen., n. sp. and the excrescences on the maxilla at the rostrum base in Choneziphius spp. are instead interpreted as areas of origin for rostral and facial muscles, acting on the nasal passages, blowhole, and melon. From a palaeobiogeographic point of view, the newly described taxa further emphasize the differences in the North Atlantic (including Iberian Peninsula) and South African Neogene ziphiid faunal lists. Even if the stratigraphic context is poorly understood, leaving open the question of the geological age for most of the dredged specimens, these differences in the composition of cold to temperate northern and southern hemisphere fossil ziphiid faunas may be explained by a warm-water equatorial barrier.
Acta Palaeontologica Polonica, Dec 1, 2009
Netherlands Journal of Geosciences - Geologie en Mijnbouw, 2015
Stable oxygen isotope values of inoceramid marine bivalve shells recovered from Bentiaba, Angola,... more Stable oxygen isotope values of inoceramid marine bivalve shells recovered from Bentiaba, Angola, are utilised as a proxy for paleotemperatures during the Late Cretaceous development of the African margin of the South Atlantic Ocean. The d 18 O values derived from inoceramids show a long-term increase from -3.2‰ in the Late Turonian to values between -0.8 and -1.8‰ in the Late Campanian. Assuming a constant oceanic d 18 O value, an ∼2‰ increase may reflect cooling of the shallow marine environment at Bentiaba by approximately 10°. Bentiaba values are offset by about +1‰ from published records for bathyal Inoceramus at Walvis Ridge. This offset in d 18 O values suggests a temperature difference of ∼5°b etween coastal and deeper water offshore Angola. Cooler temperatures implied by the d 18 O curve at Bentiaba coincide with the stratigraphic distribution of diverse marine amniotes, including mosasaurs, at Bentiaba.
Netherlands Journal of Geosciences - Geologie en Mijnbouw, 2015
ٰۣۧۙۗۢۙۗۧۙڷۣۚڷ۠ٷۢۦ۩ۣЂڷۧۘۢٷ۠ۦۙۜۨۙІ ۜۨۨۤۃҖҖ۞ۣ۩ۧ۠ٷۢۦғۗۙۛۘۦۖۡٷғۣۛۦҖІЂٰ ڷۦۣۚڷ۪ۧۙۗۦۙۧڷ۠ٷ... more ٰۣۧۙۗۢۙۗۧۙڷۣۚڷ۠ٷۢۦ۩ۣЂڷۧۘۢٷ۠ۦۙۜۨۙІ ۜۨۨۤۃҖҖ۞ۣ۩ۧ۠ٷۢۦғۗۙۛۘۦۖۡٷғۣۛۦҖІЂٰ ڷۦۣۚڷ۪ۧۙۗۦۙۧڷ۠ٷۣۢۨۘۘۆẰếẳẰẽặẬẹắẾΝẺỀẽẹẬặΝẺằΝύẰẺẾẮẴẰẹẮẰẾ ۙۦۙۜڷ۟ۗ۠Өڷۃۧۨۦۙ۠ٷڷ۠ٷۡٮ ۙۦۙۜڷ۟ۗ۠Өڷۃۣۧۢۨۤۦۗۧۖ۩ۑ ۙۦۙۜڷ۟ۗ۠Өڷۃۧۨۢۦۤۙۦڷ۠ٷۗۦۣۙۡۡӨ ۙۦۙۜڷ۟ۗ۠Өڷۃڷۙۧ۩ڷۣۚڷۧۡۦۙے ۺ۠ۦٷٮڷۙۜۨڷۣۡۦۚڷۧۢۙۡۗۙۤۧڷۦ۩ٷۣۧۧۙ۠ۤڷۘۦ۩ٷۣۧۡۧٷ۠ۙڷۣۙۢۨۗۙۢۨۧۦٷڷ۫ۙІ ۧۦ۩ٷۣۧۧۙ۠ۤڷۢڷۡۧۜۤۦۣۣۡۘۙٷۤڷۣۢڷۣۧۨۢۙۡۡۗڷۘۢٷڷٷۣ۠ۛۢۆڷۣۚڷۢٷۨۜۗۦۨۧٷٷی ۧٷۦۣیڷғۋғҒیڷۘۢٷڷ۪ۧۙ۠ٷٰۣۢçڷۣۤۡ۠íۍڷғۆڷۃۧ۩ۙۨٷیڷғۍڷۃۤ۠۩ۜۗۑڷғۑғۆڷۃۣۧۖۗٷЂڷғۋғۋڷۃۢۙۦۛۘۢۋڷЂғڷۃۢۺۣۗ۠ێڷғЂғیڷۃۣ۞úٷۦۆڷғې ڿڽڷҒڷڽڷۤۤڷۃҢڽڼھڷۺۦٷ۩ۦۖۙٯڷҖڷۙ۠ۗۨۦۆڷόẴẽẾếạẴẰỂڷҖڷٰۣۧۙۗۢۙۗۧۙڷۣۚڷ۠ٷۢۦ۩ۣЂڷۧۘۢٷ۠ۦۙۜۨۙІ Ңڽڼھڷۺۦٷ۩ۦۖۙٯڷہڽڷۃۣۙۢ۠ۢڷۘۙۜۧ۠ۖ۩ێڷۃڿۀғۀڽڼھҖۢ۞ۛғۀڽڼڽғڼڽڷۃٲۍө ھڿۀڼڼڼۀڽڿۀۀۀڿڽڼڼۑٵۨۗٷۦۨۧۖٷۛҖۦۣۘۛۙғۦۖۡٷۗ۠ۧғٷۢۦ۩ۣ۞ҖҖۃۤۨۨۜڷۃۙ۠ۗۨۦٷڷۧۜۨڷۣۨڷ۟ۢۋ ۃۙ۠ۗۨۦٷڷۧۜۨڷۙۨۗڷۣۨڷۣ۫ٱ ۫ۙІڷۧٷۦۣیڷғۋғҒیڷۘۢٷڷ۪ۧۙ۠ٷٰۣۢçڷۣۤۡ۠íۍڷғۆڷۃۧ۩ۙۨٷیڷғۍڷۃۤ۠۩ۜۗۑڷғۑғۆڷۃۣۧۖۗٷЂڷғۋғۋڷۃۢۙۦۛۘۢۋڷЂғڷۃۢۺۣۗ۠ێڷғЂғیڷۃۣ۞úٷۦۆڷғې ۡۧۜۤۦۣۣۡۘۙٷۤڷۣۢڷۣۧۨۢۙۡۡۗڷۘۢٷڷٷۣ۠ۛۢۆڷۣۚڷۢٷۨۜۗۦۨۧٷٷیڷۺ۠ۦٷٮڷۙۜۨڷۣۡۦۚڷۧۢۙۡۗۙۤۧڷۦ۩ٷۣۧۧۙ۠ۤڷۘۦ۩ٷۣۧۡۧٷ۠ۙڷۣۙۢۨۗۙۢۨۧۦٷ ڿۀғۀڽڼھҖۢ۞ۛғۀڽڼڽғڼڽۃۣۘڷҢڽڼھڷۍӨЂڷۣۢڷۙ۠ۖٷ۠ٷ۪ۆڷۃٰۣۧۙۗۢۙۗۧۙڷۣۚڷ۠ٷۢۦ۩ۣЂڷۧۘۢٷ۠ۦۙۜۨۙІڷۧғۦ۩ٷۣۧۧۙ۠ۤڷۢ ۙۦۙۜڷ۟ۗ۠Өڷۃڷۣۧۢۧۧۡۦۙێڷۨۧۙ۩ۥۙې ҢڽڼھڷۖۙٯڷۂڽڷۣۢڷھڽғھڽھғڼڿڽғہہڷۃۧۧۙۦۘۘٷڷێٲڷۃٰۛҖІЂۦۣۘۛۙғۦۖۡٷۗ۠ۧғٷۢۦ۩ۣ۞ҖҖۃۤۨۨۜڷۣۡۦۚڷۘۙۘٷۣۣ۠ۢ۫ө Netherlands Journal of Geosciences --Geologie en Mijnbouw | P age 1 of 16.
ABSTRACT Eggshells from three extant crocodilian species – Crocodylus mindorensis, Paleosuchus pa... more ABSTRACT Eggshells from three extant crocodilian species – Crocodylus mindorensis, Paleosuchus palpebrosus, and Alligator mississippiensis – has been analyzed and described. A comparison between these and other extant and fossil crocodilian eggs showed that the egg external surface ornamentation may vary between the anastomo– and the ramotuberculate (sensu Carpenter 1999) types, and the newly described rugosocavate type, characterized by an irregularly rugose surface scattered by subcircular pits that not always correspond to pore openings (Marzola et al. 2014). [...]