RMTRR OLDLIST (original) (raw)
OLDLIST, a database of old trees
OldList is a database of ancient trees. Its purpose is to identify maximum ages that different species in different localities may attain such that exceptionally old age individuals are recognized. In addition to this original OldList, Neil Pederson at Harvard Forest and colleagues at the Virginia Tech Tree Program maintain a companion Eastern OldList, focused on old trees growing in the eastern United States.
May 2019: Amazingly enough, this month has brought in two new 2000+ year-old species from two areas of the world where old trees were previously known to occur, but not quite this old. Dave Stahle and colleagues found a new stand of very old bald cypress (Taxodium distichum) from the Black River in North Carolina, USA, with a new 5th oldest known individual coming in at 2,624 years old. Dave also maintains a terrific website on their bald cypress project that old tree enthusiasts should check out. And a new paper summarizing the oldest trees in China by Jiajia Liu and colleagues has just come out, with now the 7th oldest known species to be a Qilian juniper (Juniperus przewalskii) tree that was determined to be 2230 years old.
May 2017: A new old age tree record holder was recently recognized, a Pinus longaeva (Great Basin bristlecone pine) growing in the White Mountains of eastern California. The date on this tree was reported to me by the late Tom Harlan. The tree was cored by Edmund Schulman in the summer of 1957 but unfortunately Schulman never had a chance to date the tree before his untimely death in early 1958. Starting around 2009, Harlan worked up many of the cores Schulman collected that summer of 1957, and discovered the tree's age at that time. Tom reported to me in 2010 that the tree was still alive, and the age reported to me at that time was 5062 years old. HOWEVER, as of 2017, the age of this tree has not been able to be confirmed; the core dated by Tom Harlan has not been located at the Laboratory of Tree-Ring Research. Therefore, I have removed this tree from OldList until such time as it is able to be confirmed.
Five types of tree ages are recognized in the OldList database:
- XD: crossdated
- RC: ring counted
- EX: extrapolations (usually based on ring measurements)
- HI: historic record
- C14: radiocarbon dated wood samples from a tree (added 2007)
Crossdated ages are derived through recognized dendrochronological procedures (e.g., Stokes and Smiley 1968; Swetnam, Thompson, and Sutherland 1985; Schweingruber 1987; Speer 2010). For a crossdated age, there should be no question of the age of the portion of the tree sampled, except in any portion of the ring series not confidently crossdated with either other trees at the same site or other sites in the area. Ring-counted ages are derived by simple ring counts and may contain errors in age due to missing or false rings, suppressed areas, poorly surfaced samples, or other types of tree-ring anomalies (e.g., injuries). "Age" in these first two types will invariably be a minimum age rather than true chronological age owing to the difficulty of sampling a tree exactly at the point of germination. Extrapolations are ages derived by regression from age/size relationships (e.g. Stephenson and Demetry 1995) or other mathematical or graphical methods. Ranges of ages derived by extrapolations are welcome and can be accommodated in the database. Historical ages are based on some sort of historic reference to the tree. OldList contains only a single historical age, that for a Ficus religiosa at a Buddhist Temple in Anuradhapura, Sri Lanka. A continuous account of four trees planted in the 3rd century B.C. has been kept (letter to R.J. Hartesveldt from Ambassador of Sri Lanka in the USA, December 15, 1972). An individual tree may have up to two entries in the OldList database. For example, a tree may have one entry for the age of a crossdated radial increment core and a second for an extrapolation age to a possible pith or germination date.
A note here on radiocarbon ages of potentially very old trees. There has been a lot of focus on in the media recently about very old trees that are based on radiocarbon dating of a remnant piece of wood in association with a currently living tree that is assumed to have been an ancient stem that reproduced clonally. The most recent example is "Old Tjikko", a Norway spruce (Picea abies) growing in Sweden. The living stem itself is only a few hundred years old, but there is a radiocarbon age of 9,500 years from dead wood present at its base. The living tree is argued to be only the most recent ramet of the much older individual tree genet. However, a 2016 study by G. L. Mackenthun instead argues that there is no evidence of genetic continuity between the dead and living wood portions of the tree, nor is there any evidence of clonal origination of Norway spruce in general. Thus, in the absence of any evidence of genetic continuity between dead and living portions of a stem, especially from a species otherwise not known to commonly reproduce clonally, I do not include such trees in Oldlist.
However, radiocarbon ages of trees are considered if the date came from a piece of wood that can unequivocally be associated with the individual tree itself. For example, Patrut et al. (2007) used radiocarbon dating on pieces of what may be the largest known baobab (Adansonia digitata) to determine that the tree was at least 1250± 50 years old when it recently died. This is now the oldest confirmed age of an angiosperm tree, at least twice as old as the second oldest confirmed age for an angiosperm species (although note also the addition of two oaks reported in Jones 1959 that are 866 and 930 years, sent to me by Alexei Rivera; however, we cannot find the original references and these presumably ring-counted ages cannot be confirmed). I also must at some point soon go through the recent literature on radiocarbon ages found on several tree species in South American rain forests that are in some cases several hundred to over 1000 years old. But bottom line is that confirmed (i.e., crossdated) maximum ages of conifer species are typically an order of magnitude greater than crossdated angiosperm species.
(Larger version of header photo; bristlecone pine in the Patriarch Grove in the White Mountains, California)
Notes on dates:
* Tree is still living as of late 2010s; age given is additional years since it was first sampled when this is known.
** Tree is dead; age is at time of death.
Otherwise age shown is that at time of sampling.
| Species | Age | Type | ID | Location | Collector(s), Dater(s), Reference |
|---|---|---|---|---|---|
| 4000+ years | |||||
| Pinus longaeva, Great Basin bristlecone pine | 4900** | XD | WPN-114,Prometheus | Wheeler Peak, Nevada, USA | Currey 1965; Salzer and Baisan 2018 |
| Pinus longaeva, Great Basin bristlecone pine | 4850* | XD | Methuselah | White Mountains, California, USA | Ed Schulman, Tom Harlan |
| 3000+ years | |||||
| Fitzroya cupressoides, alerce | 3622 | XD | Chile | Lara and Villalba 1993 | |
| Sequoiadendron giganteum, giant sequoia | 3266** | XD | CBR26 | Sierra Nevada, California, USA | Malcolm Hughes, Ramzi Touchan, Ed Wright |
| Sequoiadendron giganteum, giant sequoia | 3220** | XD | D-21 | Sierra Nevada, California, USA | Douglass 1919 |
| Sequoiadendron giganteum, giant sequoia | 3075** | XD | D-23 | Sierra Nevada, California, USA | Douglass 1919 |
| Sequoiadendron giganteum, giant sequoia | 3033** | XD | CMC 3 | Sierra Nevada, California, USA | Tom Swetnam, Chris Baisan |
| 2000+ years | |||||
| Juniperus occidentalis, western juniper | 2675** | XD | Scofield Juniper | Sierra Nevada, California, USA | Miles and Worthington 1998 |
| Taxodium distichum, bald cypress | 2624 | XD | BLK227 | Black River, Nort Carolina, USA | Stahle et al. 2019 |
| Pinus aristata, Rocky Mountain bristlecone pine | 2435 | XD | CB-90-11 | Central Colorado, USA | Brunstein and Yamaguchi 1992 |
| Juniperus przewalskii, Qilian juniper | 2230 | XD | Delingha, Qinghai Province, China | Liu et al. 2019 | |
| Ficus religiosa, Bodhi tree | 2217 | HI | Sri Lanka | Anonymous | |
| Sequoia sempervirens, coast redwood | 2200** | RC | Northern California, USA | Emanuel Fritz | |
| Juniperus occidentalis, western juniper | 2200 | XD/EX | Bennett Juniper | Sierra Nevada, California, USA | Peter Brown |
| Pinus balfouriana, foxtail pine | 2110 | XD | SHP7 | Sierra Nevada, California, USA | Anthony Caprio |
| 1000+ years | |||||
| Juniperus osteosperma, Utah juniper | 1967 | XD | CMU47a | Crawford Mountains, Utah, USA | Joey Pettit, Justin DeRose |
| Juniperus scopulorum, Rocky Mountain juniper | 1889 | XD | CRE 175 | Northern New Mexico, USA | Henri Grissino-Mayer, Richard Warren |
| Juniperus occidentalis, western juniper | 1810 | XD | Miles Juniper | Sierra Nevada, California, USA | Miles and Worthington 1998 |
| Pinus flexilis, limber pine | 1697 | XD | BFR-46 | Wasatch Mountains, Utah, USA | Bryan Tikalsky, Matt Bekker, Ben Bright |
| Pinus flexilis, limber pine | 1670 | XD | ERE | Northern New Mexico, USA | Tom Swetnam, Tom Harlan |
| Pinus balfouriana, foxtail pine | 1666 | XD | RCR 1 | Sierra Nevada, California, USA | Anthony Caprio |
| Pinus flexilis, limber pine | 1661 | XD | South Park, Colorado, USA | Craig Brunstein | |
| Pinus flexilis, limber pine | 1659 | XD | KET 3996 | Ketchum, Idaho, USA | Schulman 1956 |
| Thuja occidentalis, northern white cedar | 1653 | XD | FL117 | Ontario, Canada | Kelly and Larson 1997 |
| Pinus balfouriana, foxtail pine | 1649 | XD | BBL 2 | Sierra Nevada, California, USA | Anthony Caprio |
| Chamaecyparis nootkatensis, Nootka Cypress | 1636 | RC? | Vancouver Island, Canada | Les Jozsa | |
| Taxodium distichum, bald cypress | 1622 | XD | BLK 69 | Bladen County, North Carolina, USA | Stahle, Cleaveland, Hehr 1988 |
| Thuja occidentalis, northern white cedar | 1567 | XD | FL101 | Ontario, Canada | Kelly and Larson 1997 |
| Pinus flexilis, limber pine | 1542 | XD | Central Colorado, USA | Dave Yamaguchi | |
| Pinus aristata, Rocky Mountain bristlecone pine | 1438 | XD | San Francisco Peaks, Arizona, USA | Don Graybill | |
| Juniperus turkestanica, Turkestan juniper | 1437 | XD | epm0303z | Boibar Valley 3, Pakistan | J. Esper, K. Treydte, M. Winiger; Esper et al. 2009 |
| Pseudotsuga menziesii, Douglas-fir | 1350 | RC? | Vancouver Island, Canada | M. Parker and L. Jozsa (?) | |
| Juniperus occidentalis, western juniper | 1288 | XD | HL47 | Sierra Nevada, California, USA | Lisa Graumlich, MaryBeth Keifer |
| Adansonia digitata, baobab | 1275± 50 | C14 | Grootboom | Northeast Namibia | Patrut et al. 2007 |
| Pseudotsuga menziesii, Douglas-fir | 1275 | XD | BIC63 | Northern New Mexico, USA | Henri Grissino-Mayer |
| Pinus albicaulis, whitebark pine | 1267 | XD | RRR15 | Central Idaho, USA | Perkins and Swetnam 1996 |
| Juniperus tibetica, Tibetan juniper | 1265 | XD | China | Liu et al. 2019 | |
| Pinus heldreichii, Heldreich’s pine | 1229 | XD/C14 | "Oldest tree in Europe" | Pollino National Park, Italy | Piovesan et al. 2018 |
| Juniperus occidentalis, western juniper | 1172 | XD | DGS-19 | Devils Gate Summit, California, USA | Franco Biondi, Scotty Strachan |
| Pinus edulis, two-needle piñon | 1101 | XD | FMC01A | Nine Mile Canyon, Utah, USA | Chris Baisan |
| Chamaecyparis obtusa, Hinoki cypress | 1101 | XD | CIL10A | Cilan, Taiwan | Ed Wright, Ed Cook |
| Pinus albicaulis, whitebark pine | 1100+ | RC | Alberta, Canada | D. Belcher | |
| Lagarostrobus franklinii, Huon pine | 1089 | XD | Tasmania, Australia | Cook et al. 1991 | |
| Pinus albicaulis, whitebark pine | 1050 | XD | Alberta, Canada | Youngblut 2003 | |
| Pseudotsuga menziesii, Douglas-fir | 1040 | XD | ENX37 | Southern Wyoming, USA | C. Woodhouse, M. Losleben, K. Chowanski, J. Lukas, H. Adams |
| Thuja occidentalis, northern white-cedar | 1032 | XD | Ontario, Canada | Larson and Kelly 1991 | |
| Larix lyallii, subalpine larch | 1011 | XD | BAL091 | Baker Lake, Montana, USA | Malcolm Hughes, Connie Woodhouse, Peter Brown |
| < 1000 years | |||||
| Larix lyallii, subalpine larch | 980 | XD | CPL 01B 2006 | Pintler Mtns, Montana, USA | Jeremy Littell, Michael Tjoelker |
| Fokienia Hodginsii, Fujian cypress | 979 | XD | PHFH15B | Bidoup Mtn, Lam Dong Province, Vietnam | Buckley et al. 2010 |
| Pinus edulis, two-needle piñon | 973 | XD | SUN 2522 | Northeast Utah, USA | Schulman 1956 |
| Juniperus virginiana, eastern redcedar | 940 | XD | West Virginia, USA | B. Buckley, P. Krusic, E. Cook | |
| Quercus robur, English oak | 930 | RC | Lake of Bienne, Switzerland | Schadelin 1905, reported in Jones 1959 (submitted by Alexei Rivera) | |
| Pinus ponderosa, ponderosa pine | 929 | XD | Wah Wah Mtns, Utah, USA | Stan Kitchen | |
| Picea engelmannii, Engelmann spruce | 911** | XD | FCC23 | Central Colorado, USA | Brown et al. 1995 |
| Pinus monophylla, single-needle piñon | 900 | XD | MIN-08 | Pilot Range, Nevada, USA | Franco Biondi, Scotty Strachan |
| Tsuga martensiana, mountain hemlock | 889 | XD | CR54S | Glacier Bay National Park, AK, USA | Greg Wiles, Nick Wiesenberg |
| Pinus monophylla, single-needle piñon | 888 | XD | PGH-02 | Pine Grove Hills, Nevada, USA | Franco Biondi, Scotty Strachan |
| Pinus albicaulis, whitebark pine | 882 | XD | Western Alberta, Canada | Brian Luckman | |
| Pinus edulis, two-needle piñon | 878 | XD | PIC20 | Northwest Colorado, USA | Connie Woodhouse, Mark Losleben, Jeff Lukas |
| Pinus ponderosa, ponderosa pine | 877 | XD | LCW24 | NewberryCrater NM, Oregon, USA | Jim Speer, Pete Clark |
| Pseudotsuga menziesii, Douglas-fir | 870 | XD | GMX06 | Central Colorado, USA | Connie Woodhouse, Mark Losleben, Jeff Lukas |
| Pseudotsuga menziesii, Douglas-fir | 868 | XD | LAN05 | Western Colorado, USA | Connie Woodhouse, Mark Losleben, Jeff Lukas |
| Quercus petraea, sessile oak | 866 | RC? | Rankovici, Bosnia | Cited in Jones 1959 (submitted by Alexei Rivera) | |
| Pinus edulis, two-needle piñon | 859 | XD | WIL62 | Western Colorado, USA | Connie Woodhouse, Mark Losleben, Jeff Lukas |
| Pseudotsuga menziesii, Douglas-fir | 845 | XD | LAX01 | Western Colorado, USA | Connie Woodhouse, Mark Losleben, Jeff Lukas |
| Pinus ponderosa, ponderosa pine | 843 | XD | BRY4002 | Central Utah, USA | Schulman 1956 |
| Pinus ponderosa, ponderosa pine | 830 | XD | LCE89 | Newberry Crater NM, Oregon, USA | Pete Clark, K.C. Phillips |
| Larix occidentalis, western larch | 824 | XD | SL040 | Strawberry Mountains, Oregon, USA | Steve Voelker, Lara Läubli |
| Pinus ponderosa, ponderosa pine | 780 | XD | Mount Rosa, Colorado, USA | F. Brunstein | |
| Picea engelmannii, Engelmann spruce | 780 | XD | Western Alberta, Canada | Brian Luckman | |
| Larix siberica, Siberian larch | 750 | XD | OVL-5N | Ovoont, Mongolia | Baatar Nachin, Brendan Buckley, Neil Pederson |
| Pinus ponderosa, ponderosa pine | 742 | XD | Northwest Arizona, USA | Tom Harlan | |
| Larix lyalli, subalpine larch | 728 | XD | Western Alberta, Canada | Brian Luckman | |
| Pinus ponderosa, ponderosa pine | 723 | XD | Black Hills, South Dakota, USA | Bunkers et al. 1999 | |
| Tsuga mertensiana, mountain hemlock | 722 | XD | OLF14A | Cascade Mtns, Oregon, USA | Emily Heyerdahl |
| Pinus monophylla, single-needle piñon | 718 | XD | WRI-03 | White Pine Range, Nevada, USA | Franco Biondi, Scotty Strachan, Robin Tausch |
| Pinus monticola, western white pine | 707 | XD | PFT02 | Sequoia NP, California, USA | Tony Caprio |
| Pinus monophylla, single-needle piñon | 703 | XD | MTI-05 | Mt. Irish, Nevada, USA | Franco Biondi, Scotty Strachan |
| Pinus monophylla, single-needle piñon | 703 | XD | SSP-10 | Shoshone Range, Nevada, USA | Franco Biondi, Scotty Strachan |
| Pinus cembra, Swiss stone pine | 701 | XD | CALF1431 | Calimani Mts, Eastern Carpathians, Romania | Ionel Popa; Popa et al. 2009 |
| Tilia platyphyllos, bigleaf linden | 700 | HI | "Oldest tree in middle Slovakia" | Castle Bojnice, Slovakia | Miroslav Blaho; website athttp://stromy.enviroportal.sk/strom/ bojnicka-lipa?lang=en |
| Nyssa sylvatica, Tupelo | 679 | XD | New Hampshire, USA | Sperduto et al. 2000 | |
| Picea glauca, white spruce | 668 | XD | Klauane Lake, Yukon, Canada | Luckman 2003 (Brian Luckman, R. van Dorp, Don Youngblut, M. Masiokas) | |
| Abies magnifica var. shastensis, Shasta red fir | 665 | XD | CL2-5 | Klamath Mtns, California, USA | Carl Skinner, G. Everest, T. Kastner |
| Cunninghamia lanceolata, China-fir | 660 | XD | China | Liu et al. 2019 | |
| Pinus armandii, Armand pine or Chinese white pine | 634 | XD | China | Liu et al. 2019 | |
| Pinus siberica, Siberian pine | 629 | XD | TPX-16 | Tarvagatay Pass, Mongolia | Gordon Jacoby, Baatar Nachin, D. Frank |
| Pinus contorta var. murrayana, Sierra lodgepole pine | 628 | XD | UT 5G 48 | Yosemite NP, California, USA | Tony Caprio, Gus Smith, Peter Brown |
| Pinus jeffreyi, Jeffery pine | 626 | XD | TGS-02 | Truckee, California, USA | Franco Biondi, Scotty Strachan |
| Tsuga heterohylla, western hemlock | 614 | XD | YL18WHA | Glacier Bay National Park, AK, USA | Greg Wiles, Nick Wiesenberg |
| Tsuga dumosa, Himalayan hemlock | 613 | XD | China | Liu et al. 2019 | |
| Picea glauca, white spruce | 601 | XD | Aishihik Lake, Yukon, Canada | Brian Luckman, Emma Watson, & Don Youngblut 2002 | |
| Pinus strobiformis, southwestern white pine | 599 | XD | VPK02 | San Mateo Mtns, New Mexico, USA | Henri Grissino-Mayer, Jim Speer, Kiyomi Morino |
| Picea likiangensis, Lijiang spruce | 589 | XD | China | Liu et al. 2019 | |
| Tsuga canadensis, eastern hemlock | 555 | XD | Tionesta, Pennsylvania, USA | E. Cook; Cook and Cole 1991 | |
| Pinus contorta var. murrayana, Sierra lodgepole pine | 544 | XD | SGT 24 | San Bernardino Mountains, California, USA | Matt Bekker, Den Bentley, Kristin Hendrickson, Ari Menon |
| Pinus strobiformis, southwestern white pine | 538 | XD | Pinaleno Mtns, Arizona, USA | Henri Grissino-Mayer | |
| Calocedrus decurrens, incense cedar | 533 | XD | OLG02 | Sequoia NP, California, USA | Tony Caprio |
| Picea sitchensis, Sitka spruce | 532 | XD | XRSPR05 | Glacier Bay National Park, AK, USA | Greg Wiles, Nick Wiesenberg |
| Calocedrus decurrens, incense cedar | 527 | XD | CCW02 | Sequoia NP, California, USA | Tony Caprio |
| Larix sibirica, Siberian larch | 523 | XD | China | Liu et al. 2019 | |
| Picea glauca, white spruce | 522 | XD | Norton Bay, Alaska, USA | Giddings 1951 | |
| Eucalyptus regnans, mountain ash | 510±10 | XD/C14 | Styx Valley, Tasmania, Australia | Sam wood; Wood et al. 2010 | |
| Liriodendron tulipifera, yellow-poplar | 509 | Great Smoky Mountains Nat. Park, TN, USA | NADEF 2007; Dan Griffin et al. | ||
| Fagus sylvatica, European beech | 503 | XD | 1012306F | Abruzzi Nat'l Park, Italy | Piovesan et al. 2005 |
| Abies lasiocarpa, subalpine fir | 501 | XD | Southern Yukon, Canada | Luckman 2003 (Brian Luckman, M. Kenigsberg) | |
| Pinus resinosa, red pine | 500 | RC | Granite Lake, Ontario, Canada | Scott St. George | |
| Pinus bungeana, lacebark pine | 491 | XD | China | Liu et al. 2019 | |
| Abies alba, silver fir | 475 | XD | Plot 186 tree 6 | Jura Mountains, Switzerland | Badeau 1996, Bert 1993 (D. Bert, R. Schipffer, M. Becker) |
| Picea abies, Norway spruce | 468 | XD | LBG | Bavarian Forest, Germany | Rob Wilson |
| Pinus palustris, longleaf pine | 458 | XD | Weymouth Woods Nature Preserve, NC, USA | J.T. Ortegren, P. Knapp | |
| Torreya californica, California nutmeg | 455 | XD | Sierra Nevada, California, USA | Anthony Caprio | |
| Picea rubens, red spruce | 445 | RC | 05BCL901a | Fundy Escarpment, New Brunswick, Canada | B. Phillips |
| Pinus resinosa, red pine | 433 | XD | Dixon Lake Algonquin Provincial Park, Canada | Ed Cook | |
| Chamaecyparis formosensis, Formosan or Taiwan cypress | 408 | XD | Taiwan, China | Liu et al. 2019 | |
| Picea rubens, red spruce | 405 | XD | Nancy Brook, New Hampshire, USA | Paul Krusic | |
| Quercus alba, white oak | 407 | XD | Warren County, Iowa, USA | Duvick and Blasing 1983 | |
| Quercus gambelli, Gambel oak | 406 | XD | FIL383 | Chisos Mountains, New Mexico, USA | Tony Caprio |
| Quercus gambelli, Gambel oak | 401 | XD | North central Arizona, USA | Franco Biondi | |
| Quercus alba, white oak | 373 | XD | DYS08 | Belmont County, Ohio, USA | Rubino and McCarthy 2000 |
| Quercus stellata, post oak | 373 | XD | KEY13 | Osage County, Oklahoma, USA | Dave Stahle |
| Populus deltoides ssp monilifera, plains cottonwood | 370 | XD | F336 | Theo. Roosevelt NP (N unit), North Dakota, USA | Jesse and Alan Edmondson |
| Quercus rubra, red oak | 326 | XD | hem79 | Wachusett Mountain, Massachusetts, USA | Orwig et al. 2001 |
| Toxicodendron diversilobum, Pacific poison oak | 324 | RC | CG 11 2 | Aptos, Northern California, USA | Zane Moore, Steve Moore |
| Pinus echinata, shortleaf pine | 324 | XD | GMX307 | Great Smokey Mtns NP, Tennessee, USA | 2007 Dendroecological Fieldweek Fire History Group |
| Populous tremuloides, quaking aspen | 317 | XD | 1010-2-1 | Rocky Mtn NP, Colorado, USA | Dan Binkley, Peter Brown |
| Pinus echinata, shortleaf pine | 315 | XD | LAW38 | Saline County, Arkansas, USA | Dave Stahle |
| Quercus alba, white oak | 289 | XD | HT50 | Pisgah Nat'l Forest, North Carolina, USA | Speer 2001 |
| Quercus bicolor, swamp white oak | 285 | XD | RHS01a | Catkill, New York, USA | Neil Pederson, M. Hopton |
| Quercus muehlenbergii, chinquapin oak | 278 | XD | Iowa, USA | Duvick and Blasing 1983 | |
| Quercus prinus, chestnut oak | 265 | XD | GL18 | Pisgah Nat'l Forest, North Carolina, USA | Speer 2001 |
| Pinus pungens, Table Mountain pine | 232 | XD | GKA111 | Griffith Knob, Virginia, USA | Georgina DeWeese, Henri Grissino-Mayer, Charles Lafon |
| Pinus pinea, stone pine | 228 | XD | H004J | Hoyo de Pinares, Ávila, España | Fabio Natalini, Javier Vázquez-Piqué, Rafael Calama (Natalini et al. 2016) |
| Quercus lyrata, overcup oak | 218 | XD | SNA7 | Desha County Arkansas, USA | Dave Stahle |
| Populous tremuloides, quaking aspen | 216 | XD | MM-03 | Mouse Mountain Fire, Alaska, USA | David Spencer and Thomas Grant III |
| Pinus pinaster, maritime pine | 210 | XD | Pin Cazau, Landes de Gascogne Forest, France | Didier Bert, Frédéric Lagane | |
| Quercus velutina, black oak | 208 | XD | GL45 | Pisgah Nat'l Forest, North Carolina, USA | Speer 2001 |
| Xanthorrhoea preissii, balga | 200 | XD | Tree 41 | Western Australia | D. Ward |
| Quercus rubra, northern red oak | 166 | XD | GL21 | Pisgah Nat'l Forest, North Carolina, USA | Speer 2001 |
| Quercus coccinea, scarlet oak | 122 | XD | BP03 | Pisgah Nat'l Forest, North Carolina, USA | Speer 2001 |