Palaeoecology Research Papers - Academia.edu (original) (raw)

We present palaeoenvironmental, geomorphological, archaeological, and place-name data which allow a holistic assessment of the history of landscape change on Sandoy, Faroe Islands, especially in terms of the changes that occurred in... more

We present palaeoenvironmental, geomorphological, archaeological, and place-name data which allow a holistic assessment of the history of landscape change on Sandoy, Faroe Islands, especially in terms of the changes that occurred in response to the colonization of the island by humans. In contrast to other situations in the North Atlantic region, there is considerable continuity in the patterns and processes of landscape evolution across the initial

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The user has requested enhancement of the downloaded file. All in-text references underlined in blue are added to the original document and are linked to publications on ResearchGate, letting you access and read them immediately. This article appeared in a journal published by Elsevier. The attached copy is furnished to the author for internal non-commercial research and education use, including for instruction at the authors institution and sharing with colleagues.

Abstract: We present a procedure for estimating age-depth relationships in stratigraphical sequences by means of a generalized mixed-effect regression using an ancillary function for the partitioning of the fixed effect and the random... more

Abstract: We present a procedure for estimating age-depth relationships in stratigraphical sequences by means of a generalized mixed-effect regression using an ancillary function for the partitioning of the fixed effect and the random effect corresponding to the degree of representativity of the individual calibrated dates for a particular section of the sedimentary sequence. The procedure uses mid-point estimates of the calibrated ages in combination with the central distributional range as the basis for estimating the fixed relationship between age and depth. Further, it combines the variability of the calibrated age at individual layers (within-object variance) with estimation of the variability of the calibrated age distribution as a whole between the layers standardized by the fixed effect (between-object variance). These components of random variability can be considered independent, and hence the uncertainty of the estimated fixed relationship between age and depth can be estimated by combining the two random variables. The procedure follows the logic of mixed-effects models, but with prior information about the expected variance within each dated object.

Invasion ecology has made rapid progress in recent years through synergies with landscape ecology, niche theory, evolutionary ecology and the ecology of climate change. The palaeo-record of Holocene invasions provides a rich but presently... more

Invasion ecology has made rapid progress in recent years through synergies with landscape ecology, niche theory, evolutionary ecology and the ecology of climate change. The palaeo-record of Holocene invasions provides a rich but presently underexploited resource in exploring the pattern and process of invasions through time. In this paper, examples from the palaeo-literature are used to illustrate the spread of species through time and space, also revealing how interactions between invader and invaded communities change over the course of an invasion. The main issues addressed are adaptation and plant migration, ecological and evolutionary interactions through time, disturbance history and the landscape ecology of invasive spread. We consider invasions as a continuous variable, which may be influenced by different environmental or ecological variables at different stages of the invasion process, and we use palaeoecological examples to describe how ecological interactions change over the course of an invasion. Finally, the use of palaeoecological information to inform the management of invasions for biodiversity conservation is discussed.

Wesselingh, FP, Kaandorp, RJG, Vonhof, HB, Räsänen, ME, Renema, W. & Gingras, M. The nature of aquatic landscapes in the Miocene of western Amazonia: an integrated palaeontological and geochemical approach. Scripta Geologica, ...

Aim There remains some uncertainty concerning the causes of extinctions of Madagascar’s megafauna. One hypothesis is that they were caused by over-hunting by humans. A second hypothesis is that their extinction was caused by both... more

Aim There remains some uncertainty concerning the causes of extinctions of Madagascar’s megafauna. One hypothesis is that they were caused by over-hunting by humans. A second hypothesis is that their extinction was caused by both environmental change and hunting. This paper systematically addresses the second hypothesis through examination of two new pollen records from south-eastern Madagascar alongside other published records across the island.Location South-eastern Madagascar.Methods We reconstructed past vegetation and fire dynamics over the past 6000 years at two sites in south-eastern Madagascar (Ste-Luce) using fossil pollen and charcoal contained in sedimentary sequences. We investigated drivers of vegetation changes and how these, in turn, influenced faunal species in the south-east, using published climatic, archaeological and faunal records. Further, we also used published records to provide a synthesis of environmental changes on the whole island.Results Vegetation reconstructions indicate that the mosaic vegetation in the region of Ste-Luce was highly dynamic in response to climatic changes. The open woodland, surrounding the littoral forest, transformed into an ericoid grassland between c. 5800 and 5200 cal. yr bp, possibly in response to a moderate drought recorded during this period. The littoral forest was more stable between c. 5100 and 1000 cal. yr bp, with only some minor compositional changes c. 2800 cal. yr bp and between c. 1900 and 1000 cal. yr bp. Significant forest decline, however, is observed at c. 950 cal. yr bp, coinciding with a drought and a marine surge. A comparison of these results with a synthesis of published vegetation records across the island shows asynchronous vegetation changes in response to various droughts during the Holocene, except for the 950 cal. yr bp drought event, with evidence of widespread vegetation transformations and fires across the island.Main conclusions Pronounced climatic desiccation between 1200 and 700 cal. yr bp may have been the slow driver framing and triggering vegetation transformations and decline in megafaunal populations. In addition, hunting by drought-impacted human inhabitants and competition with newly introduced cattle would have amplified the impacts on megafaunal populations, leading to numerous extinctions in this period.

Multi-proxy approach was used to reconstruct the environmental conditions of remote lakes in the High Tatra Mountains (Slovakia) over the past few centuries (approximately 500-1000 years). Short sediment cores (*30 cm) taken from three... more

Multi-proxy approach was used to reconstruct the environmental conditions of remote lakes in the High Tatra Mountains (Slovakia) over the past few centuries (approximately 500-1000 years). Short sediment cores (*30 cm) taken from three morphologically similar glacial lakes distributed along altitudinal gradient (subalpine to alpine conditions) were analysed for organic matter content (LOI), diatoms and chironomids. Both descriptive and correlative approaches were used for analysing stratigraphical data. Predictive canonical correspondence analysis and co-correspondence analysis were applied to directly relate physical and biological proxies to each other. The relationship between LOI and biotic proxies was inconsistent across groups and lakes. Concordant patterns in diatom and chironomid composition were found in two non-acidified lakes. Common trends in those assemblages indicated major past environmental events such as the Little Ice Age, air pollution and lake acidification. In contrast, no relationship between the composition of diatom and chironomid assemblages was found in the formerly acidified lake, suggesting different responses of assemblages to acidification. While chironomids showed shifts that are attributable to recovery, diatoms assemblage remained relatively stable throughout the uppermost layers of the sediment record. On the other hand, climatic-driven changes in assemblages detected in the deeper layers were more pronounced in diatoms than in chironomids.

Spiral-shaped foraging trace fossils, assigned to the graphoglyptid cf. Spirorhaphe azteca, are reported from an Early Permian intertidal flat in the Robledo Mountains of southern New Mexico, USA. Remarkably similar spiral-shaped... more

Spiral-shaped foraging trace fossils, assigned to the graphoglyptid cf. Spirorhaphe azteca, are reported from an Early Permian intertidal flat in the Robledo Mountains of southern New Mexico, USA. Remarkably similar spiral-shaped structures are produced in modern intertidal flats by the paraonid polychaete Paraonis fulgens, and function as traps to capture mobile microorganisms migrating in the sediment in response to tides. We envisage a similar function for the Early Permian trace fossils. Previous studies have suggested that the lack of P. fulgens-type traces from ancient intertidal deposits indicates that such behavior only evolved geologically recently in such settings. However, this report demonstrates that such specialized foraging behavior was present in intertidal settings by at least the Early Permian. Graphoglyptids are typical of deep-marine settings, and characteristic of the Nereites ichnofacies. This represents their first undoubted occurrence in intertidal facies in the geological record. We postulate that the occurrence of graphoglyptids in deep-marine and intertidal settings is related to the predictability of resources. The scarcity of intertidal graphoglyptids in the geological record is most likely a preservational effect.

Pollen analysis has long been used as a tool to make an assessment of regional vegetation. On-site pollen samples are taken for the same purpose at some excavations, because they are often the only available contexts with good... more

Pollen analysis has long been used as a tool to make an assessment of regional vegetation. On-site pollen samples are taken for the same purpose at some excavations, because they are often the only available contexts with good preservation conditions. This paper shows that the analysis of on-and off-site pollen samples from the same region (West Frisia, the Netherlands) and time period (Dutch Bronze Age; ca. 2000-800 bc) can give very conflicting results on the type and amount of regional vegetation, in this case forest cover. Reasons for this discrepancy are discussed in relation to the use of on-site contexts by people in the past, as well as their location within a settlement. Furthermore, the potential of these contexts for the assessment of locally performed subsistence-related activities is outlined. New research directions are required to test this potential, so that wells and other man-made pollen traps can become a valuable future tool for reconstructing the past.

Kniha Počátky zemědělství ve Starém světě nabízí globální pohled na jeden z nevýznamnějších fenoménů vývoje lidstva. Nejstarší prvky zemědělství byly přítomny u lovců a sběračů, jimž je věnována značná pozornost. Na základě soudobých... more

Kniha Počátky zemědělství ve Starém světě nabízí globální
pohled na jeden z nevýznamnějších fenoménů vývoje lidstva.
Nejstarší prvky zemědělství byly přítomny u lovců a sběračů,
jimž je věnována značná pozornost. Na základě soudobých
znalostí popisuje autor dějiny řešení vzniku zemědělství
od 19. století do současnosti. Pozornost je věnována
ekologickým a společenským podmínkám, které panovaly
v přírodě a lidské společnosti na konci poslední doby ledové.
Jádrem knihy je pak líčení složitých dějů na Předním východě,
v Číně, v Africe a v Evropě tak, jak je zachycuje dnešní
bioarcheologie a paleoekologie.
Autor vidí počátky zemědělství jako dlouhodobý proces
s řadou etap, z nichž nejdůležitější bylo období pěstování
divokých rostlin na Předním východě a Číně, jež zde trvalo
řadu tisíciletí. Řada pozorování tehdejších společností
v centrech vývoje ukazuje, že sociální změny byly o něco
starší, než změny ekonomické. Pozdější plné zemědělství
charakterizuje vyšší míra sociální komplexity, vazba
na teritorium a přítomnost alespoň jedné domestikované
rostliny nebo zvířete.

In den Naturwissenschaften ist es von großem Interesse, wie sich Tierwelt, Vegetation und Landschaft am Ende der Eiszeit vor und nach dem Eingreifen des Menschen entwickelt und gegenseitig beeinflusst haben. In der Paläoökologie werden... more

In den Naturwissenschaften ist es von großem Interesse, wie sich Tierwelt, Vegetation und Landschaft am Ende der Eiszeit vor und nach dem Eingreifen des Menschen entwickelt und gegenseitig beeinflusst haben. In der Paläoökologie werden große Herbivoren der so genannten Megafauna (>45kg) als Modellorganismen genutzt, um solche komplexen Prozesse zu rekonstruieren. Die subfossilen Knochenreste dieser Arten wurden durch die häufige Bejagung in der Steinzeit im Gegensatz zu anderen Wirbeltierarten vergleichsweise repräsentativ im Erdreich abgelagert. In Mitteleuropa breiteten sich vor ca. 14000 - 13000 Jahren schon Rothirsch, Reh, Ur, Wildschwein und Elch mit unterschiedlicher Dynamik nördlich der Alpen aus. Diese Arten stellen zusammen mit dem Waldwisent und dem Wildpferd einen potenziell natürlichen Bestandteil der nacheiszeitlichen Ökosysteme Mitteleuropas dar, und durch sie induzierte Prozesse bedeuten daher 'natürliche Wildnis'. Wildpferde und Totholzkäfer, welche als Indikatoren für den Zustand der nacheiszeitlichen Urlandschaft herangezogen werden, zeigen, dass auch der Mensch spätestens seit der Jungsteinzeit einen Einfluss auf die Biodiversität hatte.

A new palaeoecological role for Archaeopteryx. (1985) R.A. Thulborn & T.L. Hamley. (1985) A new palaeoecological role for Archaeopteryx. In: The Begnnings of Birds; Proceedings of the International Archaeopteryx conference, Eichstätt,... more

A new palaeoecological role for Archaeopteryx. (1985) R.A. Thulborn & T.L. Hamley. (1985) A new palaeoecological role for Archaeopteryx. In: The Begnnings of Birds; Proceedings of the International Archaeopteryx conference, Eichstätt, 1984 (eds M.K. Hecht, J.H. Ostrom, G. Viohl & P. Wellnhofer): 81-89.

The late Quaternary history of the Lake Bolac region is reconstructed from sedimentary and pollen analysis of lake and swamp sequences and is supported by archaeological remains from lunette sediments. Freshwater deposits recovered from... more

The late Quaternary history of the Lake Bolac region is reconstructed from sedimentary and pollen analysis of lake and swamp sequences and is supported by archaeological remains from lunette sediments. Freshwater deposits recovered from lake Turangmoroke appear to date from the last glacial period, when the vegetation was composed of herbfield and grassland. Subsequent dry conditions at the end of the Pleistocene caused the deflation of some accumulated sediments. On refilling, the lake had become saline. As moisture availability increased, casuarina woodland, dominated by Allocasuarina verticillata, became the regional vegetation cover. Domination by casuarina ended ca. 8000-7000 yr BP, probably as a result of rising saline ground-water tables. Casuarinas were replaced to some degree by eucalypts, augmented by the development of a riverine Eucalyptus camaldulensis woodland. Swamp deposition commenced in the channel of neighbouring Fiery Creek at ca. 4000 yr BP, most likely as the result of a reduction in stream flow with decreased precipitation. Despite suggestions of significant impact on the vegetation by Aboriginal people from other sites in southeastern Australia, there is no evidence of any alteration here from periods of occupation dated to around 13 000 yr BP and after 3500 yr BP. The period of European occupation, though, is marked by significant changes in fire patterns, reduced tree cover, loss of floristic diversity, increased erosion rates and salinity levels, and decreased water levels.

A remarkably long period of Northern Hemispheric cooling in the 6 th century CE, which disrupted human societies across large parts of the globe, has been attributed to volcanic forcing of climate. A major tropical eruption in 540 CE is... more

A remarkably long period of Northern Hemispheric cooling in the 6 th century CE, which disrupted human societies across large parts of the globe, has been attributed to volcanic forcing of climate. A major tropical eruption in 540 CE is thought to have played a key role, but there is no consensus about the source volcano to date. Here, we present evidence for El Chichón in southern Mexico as the most likely candidate, based on a refined reconstruction of the volcano's eruption history. A new chronological framework, derived from distal tephra deposits and the world's largest Holocene beach ridge plain along the Gulf of Mexico, enabled us to positively link a major explosive event to a prominent volcanic sulfur spike in bipolar ice core records, dated at 540 CE. We speculate that voluminous tephra fall from the eruption had a severe environmental impact on Maya societies, leading to temporary cultural decline, site abandonment, and migration within the core area of Maya civilization.

Sulphur isotope data from early Archaean rocks suggest that microbes with metabolisms based on sulphur existed almost 3.5 billion years ago, leading to suggestions that the earliest microbial ecosystems were sulphur-based 1-5 . However,... more

Sulphur isotope data from early Archaean rocks suggest that microbes with metabolisms based on sulphur existed almost 3.5 billion years ago, leading to suggestions that the earliest microbial ecosystems were sulphur-based 1-5 . However, morphological evidence for these sulphur-metabolizing bacteria has been elusive. Here we report the presence of microstructures from the 3.4-billion-year-old Strelley Pool Formation in Western Australia that are associated with micrometre-sized pyrite crystals. The microstructures we identify exhibit indicators of biological affinity, including hollow cell lumens, carbonaceous cell walls enriched in nitrogen, taphonomic degradation, organization into chains and clusters, and δ 13 C values of −33 to −46 Vienna PeeDee Belemnite (VPDB). We therefore identify them as microfossils of spheroidal and ellipsoidal cells and tubular sheaths demonstrating the organization of multiple cells. The associated pyrite crystals have 33 S values between −1.65 and +1.43 and δ 34 S values ranging from −12 to +6 Vienna Canyon Diablo Troilite (VCDT) 5 . We interpret the pyrite crystals as the metabolic by-products of these cells, which would have employed sulphate-reduction and sulphur-disproportionation pathways. These microfossils are about 200 million years older than previously described 6 microfossils from Palaeoarchaean siliciclastic environments.

In the austral summer of 2006/7 the ANDRILL MIS (ANtarctic geological DRILLing-McMurdo Ice Shelf) project recovered a 1285 m sediment core from beneath the Ross Ice Shelf near Ross Island, Antarctica in a flexural moat associated with... more

In the austral summer of 2006/7 the ANDRILL MIS (ANtarctic geological DRILLing-McMurdo Ice Shelf) project
recovered a 1285 m sediment core from beneath the Ross Ice Shelf near Ross Island, Antarctica in a flexural
moat associated with volcanic loading. The upper ~600 m of this core contain sediments recording 38 glacial/
interglacial cycles of Early Pliocene to Pleistocene time, including 13 discrete diatomite units (DUs). The
longest of these, DU XI, is ~76 m-thick, and has been assigned an Early to Mid-Pliocene age (5–3 Ma). A
detailed record of the siliceous microfossil assemblages in DU XI is used in conjunction with geochemical and
sedimentological data to subdivide DU XI into four discrete subunits of continuous sedimentation. Within
each subunit, changes in diatom assemblages have been correlated with the δ18O record, providing a temporal
resolution up to 600 yr, and allowing for the construction of a detailed age model and calculation of associated
sediment accumulation rates within DU XI. Results indicate a productivity-dominated sedimentary record
with greater proportions of hemipelagic mud accumulating during relatively cool periods. This implies that
even during periods of substantial warmth, Milankovitch-paced changes in Antarctic ice volume can be linked
to ecological changes recorded in diatom assemblages.

The dodo Raphus cucullatus Linnaeus, 1758, an extinct and flightless, giant pigeon endemic to Mauritius, has fascinated people since its discovery, yet has remained surprisingly poorly known. Until the mid-19th century, almost all that... more

The dodo Raphus cucullatus Linnaeus, 1758, an extinct and flightless, giant pigeon endemic to Mauritius, has fascinated people since its discovery, yet has remained surprisingly poorly known. Until the mid-19th century, almost all that was known about the dodo was based on illustrations and written accounts by 17th century mariners, often of questionable accuracy. Furthermore, only a few fragmentary remains of dodos collected prior to the bird’s extinction exist. Our understanding of the dodo’s anatomy was substantially enhanced by the discovery in 1865 of subfossil bones in a marsh called the Mare aux Songes, situated in southeastern Mauritius. However, no contextual information was recorded during early excavation efforts, and the majority of excavated material comprised larger dodo bones, almost all of which were unassociated. Here we present a modern interdisciplinary analysis of the Mare aux Songes, a 4200-year-old multitaxic vertebrate concentration Lagerstatte. Our analysis o...

Italy is very rich in Middle Palaeolithic sites, and the Veneto region ranges among those with the best archaeozoological information. Most of the Middle Palaeolithic sites are located in caves and rock shelters situated at the mouths of... more

Italy is very rich in Middle Palaeolithic sites, and the Veneto region ranges among those with the best archaeozoological information. Most of the Middle Palaeolithic sites are located in caves and rock shelters situated at the mouths of the Alpine valleys, in the piedmont slopes. The two sites that offer the best archaeozoological data are Grotta di Fumane and Grotta S. Bernardino. Grotta S. Bernardino was occupied alternately by humans and carnivores, in particular bear. The fauna is largely dominated by ungulate remains, with red and roe deer prevalent over chamois and ibex; elk and giant deer are also present. Among the carnivores, the most frequent species is cave bear followed by lynx and leopard. Furthermore, hare, beaver and marmot are present together with remains of both fish and birds. It is possible that bears or birds of prey introduced the rare fish remains. In the Mousterian levels, hunting of the most common species was mainly directed towards young adult and adult individuals, suggesting the possibility of selective hunting. Marmot, beaver and probably bear, together with some species of birds (ducks, geese and Galliformes) were also hunted. At Grotta Fumane, Mousterian and Aurignacian levels reveal evidence of human activity related to carcass processing and bone exploitation. The most frequent ungulates are red deer, followed by roe deer and ibex; less frequent are chamois, bovids and giant deer (Megaloceros giganteus). Among the carnivores, bears (both Ursus arctos and Ursus spelaeus) are present, as are wolf, red fox and hyena (Crocuta crocuta). Hare and marmot are also present together with abundant bird remains. The most common species of bird are: Tetrao tetrix, Crex crex and Pyrrhocorax graculus. Mortality data for ungulates suggest that young adults and adults were preferentially selected. The faunal assemblage indicates that economic, ecological and climatic changes took place between the Mousterian and the Aurignacian levels.

Sphagnum peatlands in the oceanic-continental transition zone of Poland are currently influenced by climatic and anthropogenic factors that lead to peat desiccation and susceptibility to fire. Little is known about the response of... more

Sphagnum peatlands in the oceanic-continental transition zone of Poland are currently influenced by climatic and anthropogenic factors that lead to peat desiccation and susceptibility to fire. Little is known about the response of Sphagnum peatland testate amoebae (TA) to the combined effects of drought and fire. To understand the relationships between hydrology and fire dynamics, we used high-resolution multi-proxy palaeoecological data to reconstruct 2000 years of mire history in northern Poland. We employed a new approach for Polish peatlands e joint TA-based water table depth and charcoal-inferred fire activity reconstructions. In addition, the response of most abundant TA hydrological indicators to charcoal-inferred fire activity was assessed. The results show four hydrological stages of peatland development: moderately wet (from~35 BC to 800 AD), wet (from~800 to 1390 AD), dry (from~1390 to 1700 AD) and with an instable water table (from~1700 to 2012 AD). Fire activity has increased in the last millennium after constant human presence in the mire surroundings. Higher fire activity caused a rise in the water table, but later an abrupt drought appeared at the onset of the Little Ice Age. This dry phase is characterized by high ash contents and high charcoal-inferred fire activity. Fires preceded hydrological change and the response of TA to fire was indirect. Peatland drying and hydrological instability was connected with TA community changes from wet (dominance of Archerella flavum, Hyalosphenia papilio, Amphitrema wrightianum) to dry (dominance of Cryptodifflugia oviformis, Euglypha rotunda); however, no clear fire indicator species was found. Anthropogenic activities can increase peat fires and cause substantial hydrology changes. Our data suggest that increased human fire activity was one of the main factors that influenced peatland hydrology, though the mire response through hydrological changes towards drier conditions was delayed in relation to the surrounding vegetation changes.

En este ensayo contrastamos la hipótesis de “partición filogenética de nichos” para comprender los potenciales mecanismos que subyacen a las adaptaciones de los cocodrilos mesozoicos atoposáuridos bajo el modelo de la estasis coordinada.... more

En este ensayo contrastamos la hipótesis de “partición filogenética de nichos” para comprender los potenciales mecanismos que subyacen a las adaptaciones de los
cocodrilos mesozoicos atoposáuridos bajo el modelo de la estasis coordinada. Para ello exploramos factores evolutivos (filogenéticos y de diversidad) y funcionales (masa
corporal y especialización trófica), concluyendo que la evolución de este clado para el intervalo entre el Jurásico Superior y Cretácico Inferior se ha caracterizado por: a)
drástica reducción en talla, b) alta especialización ecológica. Proponemos la siguiente hipótesis: estos cocodrilos pequeños y potencialmente insectívoros ocuparon un nicho
ecológico en un momento de máxima diversificación de insectos y en ausencia de mamíferos. Los miembros del clado de los atoposáuridos se diversificaron ocuparon la
misma categoría ecomorfológica al diversificarse, lo cual retroalimentó la estabilidad del ecosistema. La pervivencia del grupo está asociada a la del ecosistema; esto sugiere que
debió darse un fuerte acoplamiento entre los ritmos de ambos.

Spiral-shaped foraging trace fossils, assigned to the graphoglyptid cf. Spirorhaphe azteca, are reported from an Early Permian intertidal flat in the Robledo Mountains of southern New Mexico, USA. Remarkably similar spiral-shaped... more

Spiral-shaped foraging trace fossils, assigned to the graphoglyptid cf. Spirorhaphe azteca, are reported from an Early Permian intertidal flat in the Robledo Mountains of southern New Mexico, USA. Remarkably similar spiral-shaped structures are produced in modern intertidal flats by the paraonid polychaete Paraonis fulgens, and function as traps to capture mobile microorganisms migrating in the sediment in response to tides. We envisage a similar function for the Early Permian trace fossils. Previous studies have suggested that the lack of P. fulgens–type traces from ancient intertidal deposits indicates that such behavior only evolved geologically recently in such settings. However, this report demonstrates that such specialized foraging behavior was present in intertidal settings by at least the Early Permian. Graphoglyptids are typical of deep-marine settings, and characteristic of the Nereites ichnofacies. This represents their first undoubted occurrence in intertidal facies in the geological record. We postulate that the occurrence of graphoglyptids in deep-marine and intertidal settings is related to the predictability of resources. The scarcity of intertidal graphoglyptids in the geological record is most likely a preservational effect.

Ecological Niche Modelling (ENM) is widely used to depict species potential occurrence according to environmental variables under different climatic scenarios. We tested the ENM approach to infer past range dynamics of cork oak, a... more

Ecological Niche Modelling (ENM) is widely used to depict species potential occurrence according to environmental variables under different climatic scenarios. We tested the ENM approach to infer past range dynamics of cork oak, a keystone species of the Mediterranean Biome, from 130 ka to the present time. Hindcasting implications would deal with a better species risk assessment and conservation management for the future. We modelled present and past occurrence of cork oak using seven ENM algorithms, starting from 63,733 spatially unique presence points at 30 arc-second resolution. Fourteen environmental variables were used and four time slices were considered (Last Interglacial, Last Glacial Maximum, mid-Holocene and present time). A threshold-independent evaluation of the goodness-of-fit of the models was evaluated by means of ROC curve and fossil or historical evidences were used to validate the results.
Four weighted average maps depicted the dynamics of area suitability for cork oak in the last 130 ka.
The derived species autoecology allowed its long-term occurrence in the Mediterranean without striking range reduction or shifting. Fossil and historical post-processing validation support the modelled past spatial extension and a neglected species presence at Levantine until the recent time.
Despite the severe climatic oscillation since the Last Glacial Maximum, cork oak potential distribution area experienced limited range changes, confirming its strong link with the Mediterranean Basin. The ecological amplitude of Quercus suber could be therefore adopted as a reference to trace the Mediterranean bioclimate area. A better knowledge of the past events of Mediterranean vegetation, a wider range of study species and environmental determinants are essential to inform us about its current state, its sensitivity to human impact and the potential responses to future changes.

The article presents the diagnosis and aetiology of an arthropathological horse (Equus sp.) radius from the Middle Palaeolithic site of Pfeddersheim (Rhine-Hesse, Germany). For the contextualisation of the results the chronological and... more

The article presents the diagnosis and aetiology of an arthropathological horse (Equus sp.) radius from the Middle Palaeolithic site of Pfeddersheim (Rhine-Hesse, Germany). For the contextualisation of the results the chronological and archaeological frame of the site, the taphonomic history of the faunal assemblage and site formation are summarised. A fracture of the ulna caused a malposition of the humeroradial joint, that developed into a degenerative arthrosis with severe grooving of the articular surface of the radius, extension of the joint by new bone formation and exostosis around the epiphysis. The progression of the disease from the trauma to the final grade of arthrosis and how it affected the horse's life is portrayed. Eventually, the horse fell prey to a large carnivore. Finally, the palaeoecological significance of this very specific palaeopathological finding is discussed.

The diatom stratigraphy of a 23-m sediment core from Lake Tilo, a maar lake in the Ethiopian Rift Valley, provides a 10,000-yr record of lake salinity and trophic status. Until 5500 14 C yr B.P., the phytoplankton was dominated by... more

The diatom stratigraphy of a 23-m sediment core from Lake Tilo, a maar lake in the Ethiopian Rift Valley, provides a 10,000-yr record of lake salinity and trophic status. Until 5500 14 C yr B.P., the phytoplankton was dominated by Aulacoseira granulata, with only minor changes in the abundance of other diatoms; the lake was over 50 m deep, eutrophic, and oligosaline. At 5500 yr B.P., geothermal groundwater inflow, inferred from calcite and silica deposition rates, declined abruptly, and the lake became more oligotrophic, as indicated by a rapid rise in Cyclotella stelligera. About 4500 yr B.P., lake salinity began to increase, reaching approximately its present state ca. 2500 yr B.P., but with a temporary reversal to lower salinity at 4000 -3500 yr B.P. The record shows no evidence of salinity increases equivalent to early Holocene low stands of the larger river-fed Rift Valley lakes, probably because of high rates of geothermal groundwater influx. It responded to reduced rainfall at 4500 yr B.P., when levels of the larger lakes also fell, because geothermal groundwater flux had diminished 1000 years earlier, independently of climate. Because geothermal groundwaters can form a significant proportion of a crater lake's hydrological budget and dominate its salinity budget, these results show that variable geothermal groundwater flux can override lake hydrochemical response to climate change. Palaeoclimatic interpretation of palaeosalinity proxies from the sediments of volcanic crater lakes should be approached with caution. © 1999 University of Washington.

An assemblage of the shallow-water radiolitid Eoradiolites liratus (Conrad, 1852) is described from the Upper Cenomanian Galala Formation at Saint Paul (southern Galala, Eastern Desert, Egypt). At this locality, the stratigraphically... more

An assemblage of the shallow-water radiolitid Eoradiolites liratus (Conrad, 1852) is described from the Upper Cenomanian Galala Formation at Saint Paul (southern Galala, Eastern Desert, Egypt). At this locality, the stratigraphically youngest rudists occur just below the latest Cenomanian ammonite Vascoceras cauvini/V. rumeaui Biozone. Palaeogeographically, E. liratus shows affinities with southern parts of the Mediterranean Tethyan Realm and a close relationship with the Apulian Plate in particular. The species is restricted to the Cenomanian of the Eastern Desert, whereas in Sinai, as well as in the adjacent eastern Mediterranean and other countries in the Near East, it occurs in older (Albian) deposits. The present collection shows a wide range of variation in shape and ornament, yet this does not involve large differences in individual size. The variation within this assemblage of E. liratus is mainly ecophenotypic, rather than ontogenetic. Bioerosion traces are common in the collection available; the number of borings is high and they are concentrated mainly in the commissural area, in radial bands and interbands as well as in left valves. Such borings indicate a fairly long period of post-mortem exposure for these shells.

The harp seal (Pagophilus groenlandicus), today a subarctic species with breeding populations in the White Sea, around the Jan Mayen Islands and Newfoundland, was a common pinniped species in the Baltic Sea during the mid-and late... more

The harp seal (Pagophilus groenlandicus), today a subarctic species with breeding populations in the White Sea, around the Jan Mayen Islands and Newfoundland, was a common pinniped species in the Baltic Sea during the mid-and late Holocene. It is puzzling how an ice dependent species could breed in the Baltic Sea during the Holocene Thermal Maximum (HTM), and it remains unclear for how long harp seals bred in the Baltic Sea and when the population became extirpated. We combined radiocarbon dating of harp seal bones with zooarchaeological, palaeoenvironmental and stable isotope data to reconstruct the harp seal occurrence in the Baltic Sea. Our study revealed two phases of harp seal presence and verifies that the first colonization and establishment of a local breeding population occurred within the HTM. We suggest that periods with very warm summers but cold winters allowed harp seals to breed on the ice. Human pressure, salinity fluctuations with consequent changes in prey availability and competition for food resources, mainly cod, resulted in physiological stress that ultimately led to a population decline and local extirpation during the first phase. The population reappeared after a long hiatus. Final extinction of the Baltic Sea harp seal coincided with the Medieval Warm Period. Our data provide insights for the first time on the combined effects of past climatic and environmental change and human pressure on seal populations and can contribute with new knowledge on ongoing discussions concerning the impacts of such effects on current arctic seal populations.

This paper presents stratigraphical details of the Chedworth Nature Reserve Road Bridge Cutting, a detailed analysis of the fossils of one particular bed within the Clypeus Grit Member and a comparison of this assemblage with a more... more

This paper presents stratigraphical details of the Chedworth Nature Reserve Road Bridge Cutting, a detailed analysis of the fossils of one particular bed within the Clypeus Grit Member and a comparison of this assemblage with a more typical Clypeus Grit bed on Cleeve Hill. The Chedworth CG Soft Bed has very few deep-burrowing bivalves, a lower proportion of encrusting animals and a higher proportion of crushed Stiphrothyris shells. The latter two features could indicate a more rapid sedimentation rate for the bed.

Palaeobotanical specimens in private or public historical collections include important fossils, and modern studies involving taxonomic revisions can renew the scientific value of forgotten records. In Italy, information on past floras... more

Palaeobotanical specimens in private or public historical collections include important fossils, and modern studies involving taxonomic revisions can renew the scientific value of forgotten records. In Italy, information on past floras can be also obtained through the re-examination of the published drawings and pictures, or survey of the original collections. This paper presents the taxonomic revision of the macrofossils in the palaeobotanical collection, originally from San Venanzio (Modena), collected by Francesco Coppi at the end of the 19th century. Fossils were taken from the Argille Azzurre formation (Lower Pliocene-Lower Pleistocene). Re-sampling was not carried out because the sampling point at present lies under the houses of a village, increasing the importance of the museological collection. Descriptions of 76 fossil leaves are reported. Sporomorphs were extracted from the clayey limestone where three macrofossils had fossilized. Though only scattered information on the past plant cover has been randomly preserved, the plant macro-and microfossils provide a contribution to the research on the Plio-Pleistocene flora of Northern Apennines, and add information which can be useful to improve knowledge on local palaeoenvironmental reconstructions. Animal and plant fossils were deposited in a coastal or delta area. Pollen flora mainly reflected the vegetation distributed in the highest belts, far from the site of deposition, while macrofossils prevalently reflected local vegetation.

Diprotodon Owen, was one of the first fossil mammals described from Australia and has the distinction of being the largest ever marsupial. However, until recently its taxonomy was unclear and knowledge of its continental distribution,... more

Diprotodon Owen, was one of the first fossil mammals described from Australia and has the distinction of being the largest ever marsupial. However, until recently its taxonomy was unclear and knowledge of its continental distribution, palaeoecology and evolution was poorly known. This paper describes cranial elements from an unusually small-bodied Diprotodon from the Early Pleistocene Nelson Bay Formation, Portland, Australia. It is intermediate in size between the smaller-bodied Pliocene ancestor, Euryzygoma dunense de Vis 1895 and the larger-bodied Late Pleistocene D. optatum Owen, . However, it is morphologically inseparable from Late Pleistocene Diprotodon and is here referred to as D. ?optatum. A temporal morphocline most parsimoniously explains the medium body size of the Nelson Bay taxon; thus we provide the first evidence of a transitional form within Diprotodon. The cause(s) of gigantism of the E. dunense-D. ?optatum-D. optatum lineage through the late Cainozoic is unclear but most likely involved the physiological response to changes in the physical and/or biotic environment. Although the size changes within the lineage may have been initially advantageous, they were most likely disadvantageous when the optimum body size was obtained. Our results suggest that Diprotodon was at its largest ever body size during the Late Pleistocene - the time period that coincided with its extinction.

Abstract: Although their record extends back to the Early Ordovician, the occurrence of fossil starfish (Echinodermata: Asteroidea) is dependent almost exclusively upon horizons of exceptional preservation. Thus, asteroids found in... more

Abstract: Although their record extends back to the Early Ordovician, the occurrence of fossil starfish (Echinodermata: Asteroidea) is dependent almost exclusively upon horizons of exceptional preservation. Thus, asteroids found in Silurian obrution deposits of the English Midlands and Welsh Borderlands are particularly significant to an understanding of the early diversity of the group. Six species are described here: Hudsonaster? carectum sp. nov. (Hudsonasteridae), from the lower part of the Lower Elton Formation; and, from the Much Wenlock Limestone Formation, the hudsonasterids Doliaster brachyactis gen. et sp. nov. and Siluraster? ketleyi (Spencer, 1916), the lepidasterids Lepidaster grayi Forbes, 1850 and Lepidactis wenlocki Spencer, 1918, and the palasterinid Palasterina orchilocalia sp. nov. Though few in number, they show a diverse range of body morphologies when compared with Ordovician taxa: L. wenlocki had long, slender rays when fully grown whereas D. brachyactis is the first asteroid with the short-rayed body form of extant cushion stars. Most distinctive of all is L. grayi, the earliest multiradiate taxon known, all complete specimens of which have 13 rays. This morphological variety is interpreted as indicating that by the Early Silurian starfish were exploiting a wide range of feeding habits and ecological niches.

Although Cretaceous fossils (coal excluded) from Victoria, Australia, were first reported in the 1850s, it was not until the 1950s that detailed studies of these fossils were undertaken. Numerous fossil localities have been identified in... more

Although Cretaceous fossils (coal excluded) from Victoria, Australia, were first reported in the 1850s, it was not until the 1950s that detailed studies of these fossils were undertaken. Numerous fossil localities have been identified in Victoria since the 1960s, including the Koonwarra Fossil Bed (Strzelecki Group) near Leongatha, the Dinosaur Cove and Eric the Red West sites (Otway Group) at Cape Otway, and the Flat Rocks site
(Strzelecki Group) near Cape Paterson. Systematic exploration over the past five decades has resulted in the collection of thousands of fossils representing various plants, invertebrates and vertebrates. Some of the best-preserved and most diverse Hauterivian–Barremian floral assemblages in Australia derive from outcrops of the lower Strzelecki Group in the Gippsland Basin. The slightly younger Koonwarra Fossil Bed (Aptian) is a Konservat-Lagerstätte that also preserves abundant plants, including one of the oldest known flowers. In addition, insects, crustaceans (including the only syncaridans known from Australia between the Triassic and the present), arachnids (including Australia’s only known opilione), the stratigraphically youngest xiphosurans from Australia, bryozoans, unionoid molluscs and a rich assemblage of actinopterygian fish are known from the Koonwarra Fossil Bed. The oldest known—and only Mesozoic—fossil feathers from the Australian continent constitute the only evidence for tetrapods at Koonwarra. By contrast, the Barremian–Aptian-aged deposits at the Flat Rocks site, and the Aptian–Albian-aged strata at the Dinosaur Cove and Eric the Red West sites, are all dominated by tetrapod fossils, with actinopterygians and dipnoans relatively rare. Small ornithopod (=basal neornithischian) dinosaurs are numerically common, known from four partial skeletons and a multitude of isolated bones. Aquatic meiolaniform turtles constitute another prominent faunal element, represented by numerous isolated bones and articulated carapaces and plastrons. More
than 50 specimens—mostly lower jaws—evince a high diversity of mammals, including monotremes, a multituberculate and several enigmatic ausktribosphenids. Relatively minor components of these fossil assemblages are diverse theropods (including birds), rare ankylosaurs and ceratopsians, pterosaurs, non-marine plesiosaurs and a lepidosaur. In the older strata of the upper Strzelecki Group, temnospondyl amphibians—the youngest known worldwide—are a conspicuous component of the fauna, whereas crocodylomorphs appear to be present only in up-sequence deposits of the Otway Group. Invertebrates are uncommon, although decapod crustaceans and unionoid bivalves have been described. Collectively, the Early Cretaceous biota of Victoria provides insights into a unique Mesozoic high-latitude palaeoenvironment and elucidates both palaeoclimatic and palaeobiogeographic changes throughout more than 25 million years of geological time.

How have humans colonised the entire planet and reshaped its ecosystems in the process? This unique and groundbreaking collection of essays explores human movement through time, the impacts of these movements on landscapes and other... more

How have humans colonised the entire planet and reshaped its ecosystems in the process? This unique and groundbreaking collection of essays explores human movement through time, the impacts of these movements on landscapes and other species, and the ways in which species have co-evolved and transformed each other as a result. Exploring the spread of people, plants, animals, and diseases through processes of migration, colonisation, trade, and travel, it assembles a broad array of case studies from the Pliocene to the present. The contributors from disciplines across the humanities and natural sciences are senior or established scholars in the fields of human evolution, archaeology, history, and geography.

Talk on submerged forests in the UK, looking at Orkney, Severn Estuary and the Pett Level.

The hypothesis that the Ediacara biota were giant protozoans is tested by considering the external morphology, internal organization, suggested fossil representatives and molecular phylogeny of the xenophyophores. From this analysis, we... more

The hypothesis that the Ediacara biota were giant protozoans is tested by considering the external morphology, internal organization, suggested fossil representatives and molecular phylogeny of the xenophyophores. From this analysis, we find no case to support a direct relationship. Rather, the xenophyophores are here regarded as a group of recently evolved Foraminifera and are hence unlikely to have a record from the Ediacaran Period. Further from the growth dynamics of Foraminifera, they are also unlikely to be related to the Palaeopascichnus organism. We also find significant distinctions in the growth dynamics of Palaeopascichnus and organ-isms usually referred to the Ediacara biota, such as Charnia and Dickinsonia. Developmental analysis of the Palaeopascichnus -central to the xenophyophore hypothesis -reveals unusual, protozoan features, including evidence for chaotic repair structures, for mergence of coeval forms, as well as complex bifurcations. These observations suggest that Palaeopascichnus is a body fossil of an unidentified protozoan but is unrepresentative of Ediacaran body construction, in general.